Cranial morphology of Heckerochelys romani Sukhanov, 2006, a stem turtle from the Middle Jurassic of European Russia, with implications for the paleoecology of stem turtles

LITERATURE CITED

• Alekseev, A. S., Barskov, I.S., Gordenko, N.V., Krasnikov, N.M., Rozanova, A.A., Smirnova, S.B., Fokin, P.A., Agadzhanyan, A.K., Efimov, M.B., Kabanov, P.B. & Krasilov, V.A. (2001). Discovery of unique Middle Jurassic fauna and flora in the Moscow suburbs. Doklady Earth Sciences 377(3), 359–362. [in Russian]
   Web of Science ®Google Scholar
• Alifanov V. R. & Sennikov A. G. (2001). About discovery of dinosaur remains in Moscow region. Doklady Earth Sciences 376(1), 73–75. [in Russian]
   Web of Science ®Google Scholar
• Anquetin, J. (2010). The anatomy of the basal turtle Eileanchelys waldmani from the Middle Jurassic of the Isle of Skye, Scotland. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 101(1), 67–96.
   Google Scholar
• Anquetin, J. (2011). Evolution and palaeoecology of early turtles: a review based on recent discoveries in the Middle Jurassic. Bulletin de la Société géologique de France, 182(3), 231–240.
   Google Scholar
• Anquetin, J. (2012). Reassessment of the phylogenetic interrelationships of basal turtles (Testudinata). Journal of Systematic Palaeontology, 10(1), 3–45.
   Web of Science ®Google Scholar
• Anquetin, J., Barrett, P. M., Jones, M. E., Moore-Fay, S., & Evans, S. E. (2009). A new stem turtle from the Middle Jurassic of Scotland: new insights into the evolution and palaeoecology of basal turtles. Proceedings of the Royal Society B: Biological Sciences, 276(1658), 879–886.
   PubMed Web of Science ®Google Scholar
• Averianov, A. O. (2002). Review of Mesozoic and Cenozoic sea turtles from the former USSR. Russian Journal of Herpetology, 9(2), 137–154.
   Google Scholar
• Averianov, A. O., & Zverkov, N. G. (2020). New diplodocoid sauropod dinosaur material from the Middle Jurassic of European Russia. Acta Palaeontologica Polonica, 65(3), 499–509.
   Web of Science ®Google Scholar
• Baur, G. (1887). Osteologische Notizen über Reptilien (Fortsetzung II). Zoologischer Anzeiger, 10, 241–268.
   Google Scholar
• Benson, R. B., Domokos, G., Várkonyi, P. L., & Reisz, R. R. (2011). Shell geometry and habitat determination in extinct and extant turtles (Reptilia: Testudinata). Paleobiology, 37(4), 547–562.
   Web of Science ®Google Scholar
• Bramble D. M., & Wake D. B. (1985). Feeding mechanisms of lower tetrapods. In M. Hildebrand, D. M. Bramble, K. F. Liem, and D. Wake (Eds.), Functional Vertebrate Morphology (pp. 230–261). Harvard Univ. Press, Cambridge, MA.
   Google Scholar
• Carpenter, K., & Bakker, R. T. (1990). A new latest Jurassic vertebrate fauna, from the highest levels of the Morrison Formation at Como Bluff, Wyoming. Part II. A new baenid turtle. Hunteria, 2(6), 1–19.
   Google Scholar
• Cherepanov, G. O. (2019). Morphogenetic and constructional differences of the carapace of aquatic and terrestrial turtles and their evolutionary significance. Journal of Morphology, 280(10), 1571–1581.
   PubMed Web of Science ®Google Scholar
• Cuvier, G. (1825). Recherches sur les ossemens fossiles (Vol. 5). Dufour-D'Ocagne.
   Google Scholar
• Danilov, I. G. (2005). Die fossilen Schildkröten Europas. In U. Fritz (Ed.), Handbuch der Reptilien und Amphibien Europas. Band 3/IIIB: Schildkröten (Testudines) II (Cheloniidae, Dermochelyidae, Fossile Schildkröten Europas (pp. 329–441). Wiebelsheim, Aula-Verlag.
   Google Scholar
• Danilov, I. G., & Obraztsova, E. M. (2015). Taxonomic position of Oxemys gutta Nessov, 1977, a problematic turtle from the Upper Cretaceous of Uzbekistan, and basisphenoid morphology in some groups of Late Mesozoic turtles of Asia. Paleontological Journal, 49(3), 279–292.
   Web of Science ®Google Scholar
• Danilov, I. G., Syromyatnikova, E. V., & Sukhanov, V. B. (2017). Subclass Testudinata. In E. I. Vorobyeva, A. V. Lopatin, and M. A. Shishkin (Eds.), Fossil vertebrates of Russia and adjacent countries, Fossil reptiles and birds, Part 4 (pp. 27–343). Moscow: GEOS. [in Russian]
   Google Scholar
• Dziomber, L., Joyce, W. G., & Foth, C. (2020). The ecomorphology of the shell of extant turtles and its applications for fossil turtles. PeerJ, 8, e10490.
   PubMed Web of Science ®Google Scholar
• Dudgeon, T. W., Livius, M. C., Alfonso, N., Tessier, S., & Mallon, J. C. (2021). A new model of forelimb ecomorphology for predicting the ancient habitats of fossil turtles. Ecology and Evolution, 11(23), 17071–17079.
   PubMed Web of Science ®Google Scholar
• Evers, S. W., & Joyce, W. G. (2020). A re-description of Sandownia harrisi (Testudinata: Sandownidae) from the Aptian of the Isle of Wight based on computed tomography scans. Royal Society Open Science, 7(2), 191936.
   PubMed Web of Science ®Google Scholar
• Evers, S. W., Rollot, Y., & Joyce, W. G. (2020). Cranial osteology of the Early Cretaceous turtle Pleurosternon bullockii (Paracryptodira: Pleurosternidae). PeerJ, 8, e9454.
   PubMed Web of Science ®Google Scholar
• Evers, S. W., Ponstein, J., Jansen, M. A., Gray, J. A., & Fröbisch, J. (2022). A systematic compendium of turtle mandibular anatomy using digital dissections of soft tissue and osteology. The Anatomical Record, 306, 1228–1303.
   Google Scholar
• Foth, C., Rabi, M., & Joyce, W. G. (2017). Skull shape variation in extant and extinct Testudinata and its relation to habitat and feeding ecology. Acta Zoologica, 98(3), 310–325.
   Web of Science ®Google Scholar
• Gaffney, E. S. (1972). An illustrated glossary of turtle skull nomenclature. American Museum Novitates, 2486, 1–33.
   Google Scholar
• Gaffney, E. S. (1975). A taxonomic revision of the Jurassic turtles Portlandemys and Plesiochelys. American Museum Novitates, 2574, 1–19.
   Google Scholar
• Gaffney, E. S. (1979). Comparative cranial morphology of Recent and fossil turtles. Bulletin of the American Museum of Natural History, 164, 69–376.
   Google Scholar
• Gaffney, E. S. (1983). The cranial morphology of the extinct horned turtle, Meiolania platyceps, from the Pleistocene of Lord Howe Island, Australia. Bulletin of the American Museum of Natural History, 175, 361–480.
   Web of Science ®Google Scholar
• Gaffney, E. S. (1990). The comparative osteology of the Triassic turtle Proganochelys. Bulletin of the American Museum of Natural History, 194, 1–263.
   Google Scholar
• Gaffney, E. S., & Jenkins Jr., F. A. (2010). The cranial morphology of Kayentachelys, an Early Jurassic cryptodire, and the early history of turtles. Acta Zoologica, 91(3), 335–368.
   Web of Science ®Google Scholar
• Gaffney, E. S., & Kitching, J. W. (1994). The most ancient African turtle. Nature, 369, 55–58.
   Web of Science ®Google Scholar
• Gaffney, E. S., Hutchison, J. H., Jenkins Jr, F. A., & Meeker, L. J. (1987). Modern turtle origins: the oldest known cryptodire. Science, 237(4812), 289–291.
   PubMed Web of Science ®Google Scholar
• Gaffney, E. S., Rich, T. H., Vickers-Rich, P., Constantine, A., Vacca, R., & Kool, L. (2007). Chubutemys, a new eucryptodiran turtle from the Early Cretaceous of Argentina, and the relationships of the Meiolaniidae. American Museum Novitates, 2007(3599), 1–35.
   Google Scholar
• Gordenko, N. V. (2008). Middle Jurassic flora of the Peski locality (Moscow Region): Systematics, paleoecology, and phytogeography. Paleontological Journal, 42(12), 1285–1382.
   Web of Science ®Google Scholar
• Grandidier, A. (1867). Liste des reptiles nouveaux découverts, en 1866, sur la côte sud-ouest de Madagascar. Revue et Magazine de Zoologie (Paris), Sér. 2, 19, 232–234
   Google Scholar
• Gray, J. E. (1830). A synopsis of the species of Class Reptilia. In Griffith, E & E. Pidgeon: The animal kingdom arranged in conformity with its organisation by the Baron Cuvier with additional descriptions of all the species hither named, and of many before noticed Whittaker, Treacher and Co., London.
   Google Scholar
• Hay, O. P. (1905). On the group of fossil turtles known as the Amphichelydia: with remarks on the origin and relationships of the suborders, superfamilies, and families of Testudines. Bulletin of American Museum of Natural History, 21, 137–175.
   Google Scholar
• Hermanson, G., Benson, R. B., Farina, B. M., Ferreira, G. S., Langer, M. C., & Evers, S. W. (2022). Cranial ecomorphology of turtles and neck retraction as a possible trigger of ecological diversification. Evolution, 76(11), 2566–2586.
   PubMed Web of Science ®Google Scholar
• Joyce, W. G., Parham, J. F., & Gauthier J. A. (2004). Developing a protocol for the conversion of rank-based taxon names to phylogenetically defined clade names, as exemplified by turtles. Journal of Paleontology, 78(5), 989–1013.
   Web of Science ®Google Scholar
• Joyce, W. G. (2007). Phylogenetic relationships of Mesozoic turtles. Bulletin of the Peabody Museum of Natural History, 48(1), 3–102.
   Google Scholar
• Joyce, W. G. (2017). A review of the fossil record of basal Mesozoic turtles. Bulletin of the Peabody Museum of Natural History, 58(1), 65–113.
   Web of Science ®Google Scholar
• Joyce, W. G., Parham, J. F., Anquetin, J., Claude, J., Danilov, I. G., Iverson, J. B., Kear, B., Lyson, T. R., Rabi, M., & Sterli, J. (2020). Testudinata. In K. de Queiroz, P. D. Cantino, & J. A. Gauthier (Eds.), Phylonyms – A Companion to the PhyloCode (pp. 1045–1048). CRC Press: Boca Raton.
   Google Scholar
• Joyce, W. G., Anquetin, J., Cadena, E. A., Claude, J., Danilov, I. G., Evers, S. W., Ferreira, G. S., Gentry, A. D., Georgalis, G. L., Lyson, T. R., & Pérez-García, A. (2021a). A nomenclature for fossil and living turtles using phylogenetically defined clade names. Swiss Journal of Palaeontology, 140(1), 1–45.
   Web of Science ®Google Scholar
• Joyce, W. G., Mäuser, M., & Evers, S. W. (2021b). Two turtles with soft tissue preservation from the platy limestones of Germany provide evidence for marine flipper adaptations in Late Jurassic thalassochelydians. PLoS ONE, 16(6), e0252355.
   PubMed Web of Science ®Google Scholar
• Joyce, W. G., & Gauthier, J. A. (2004). Palaeoecology of Triassic stem turtles sheds new light on turtle origins. Proceedings of the Royal Society of London. Series B: Biological Sciences, 271(1534), 1–5.
   PubMed Web of Science ®Google Scholar
• Joyce, W. G., Rabi, M., Clark, J. M., & Xu, X. (2016). A toothed turtle from the Late Jurassic of China and the global biogeographic history of turtles. BMC Evolutionary Biology, 16(1), 1–29.
   PubMedGoogle Scholar
• Khosatzky, L. I. (1998). Big turtle of the Late Cretaceous of Mongolia. Russian Journal of Herpetology, 4(2), 148–154.
   Google Scholar
• Klein, I. T. (1760). Klassification und kurze Geschichte der Vierfüßigen Thiere (translation by F. D. Behn). Jonas Schmidt, Lübeck.
   Google Scholar
• Krupina, N. I. (1995). The first finding of Jurassic ceratodontids (Dipnoi) from the Moscow Region. Paleontological Journal, 1, 129–131. [in Russian].
   Google Scholar
• Latreille, P. A. (1801). In C.N.S. Sonnini de Manoncourt and P.A. Latreille. Histoire Naturelle des Reptiles, avec Figures Déssinnées d'après Nature. Vol. 1. Détérville, Paris, xx + 280 p.
   Google Scholar
• Le Conte, J. (1830). Description of the species of North American tortoises. Annals of the Lyceum of natural History of New York, 3(1), 91–131.
   Google Scholar
• Lichtig, A. J., & Lucas, S. G. (2017). A simple method for inferring habitats of extinct turtles. Palaeoworld, 26(3), 581–588.
   Web of Science ®Google Scholar
• Linnaeus, C. (1758). Systema naturae (Vol. 1, p. 824). Laurentii Salvii: Stockholm.
   Google Scholar
• Meylan, P. A., Moody, R. T., Walker, C. A., & Chapman, S. D. (2000). Sandownia harrisi, a highly derived trionychoid turtle (Testudines: Cryptodira) from the Early Cretaceous of the Isle of Wight, England. Journal of Vertebrate Paleontology, 20(3), 522–532.
   Web of Science ®Google Scholar
• Northmore, D. P. M., & Granda, A. M. (1991). Ocular dimensions and schematic eyes of freshwater and sea turtles. Visual neuroscience, 7(6), 627–635.
   PubMed Web of Science ®Google Scholar
• Novikov, I. V., Lebedev, O. A., & Alifanov, V. R. (1998). New Mesozoic vertebrate fossil sites of Russia. In J. W. M. Jagt, P. H. Lambers, E. W. A. Mulder, & A. S. Schulp (Eds.), Third European Workshop on Vertebrate Palaeontology (pp. 6–9). Maastricht.
   Google Scholar
• Obraztsova, E. M. (2021). Evolutionary trends and homology of basicranial structures in Mesozoic turtles. In Turtle Evolution Symposium 2021, Trelew, Argentina, Book of abstracts (p. R96). PE-APA.
   Google Scholar
• Obraztsova, E. M. (2022a). Drawings in the morphological description: theoretical principles and methods, on the example of preparing an osteological illustration. In K. G. Mikhailov (Ed.), Evolutionary and functional morphology of Vertebrates (pp. 234–236). Moscow, KMK Scientific Press. [in Russian]
   Google Scholar
• Obraztsova, E. M. (2022b). Morphofunctional features and evolutionary transformations of the turtle ear. In K. G. Mikhailov (Ed.), Evolutionary and functional morphology of Vertebrates (pp. 234–236). Moscow, KMK Scientific Press. [in Russian]
   Google Scholar
• Obraztsova, E. M. (2022c). The process of morphological evolution as an expansion of the “space of characters” in time. In K. G. Mikhailov (Ed.), Evolutionary and functional morphology of Vertebrates (pp. 234–236). Moscow, KMK Scientific Press. [in Russian]
   Google Scholar
• Obraztsova, E. M., Danilov I. G., & Sukhanov V. B. (2018). Comments on the cranial morphology of Heckerochelys romani Sukhanov, 2006, a basal turtle from the Middle Jurassic of European Russia. In R. Hirayama, S. Kuratani, A. Takahashi, Y. Nakajima, T. Sonoda, Y. Nishioka, & M. Yoshida (Eds.), Turtle Evolution Symposium 2018 Tokyo (pp. 69–72). Tokyo: Scidinge Hall Verlag, Tübingen.
   Google Scholar
• Obraztsova, E. M., Krasnolutskii S. A., Sukhanov V. B., & Danilov I. G. (2022). Xinjiangchelyid turtles from the Middle Jurassic of the Berezovsk coal mine (Krasnoyarsk Territory, Russia): systematics, skeletal morphology, variation, relationships and palaeobiogeographic implications. Journal of Systematic Palaeontology, 20(1), 1–61.
   Web of Science ®Google Scholar
• Owen, R. (1842). Report on British fossil reptiles. Part II. In Report of the Eleventh Meeting of the British Association for the Advancement of Science, 1842 (pp. 60–204), London.
   Google Scholar
• Owen, R. (1886). Description of Fossil Remains of Two Species of a Megalanian Genus (Meiolania) from “Lord Howe's Island”. Philosophical Transactions of the Royal Society of London, 177, 471–480.
   Google Scholar
• Pashchenko, D. I., Kuzmin, I. T., Sennikov, A. G., Skutschas, P. P., & Efimov, M. B. (2018). On the finding of neosuchians (Neosuchia, Crocodyliformes) in the Middle Jurassic (Bathonian) deposits of the Moscow Region. Paleontological Journal, 52(5), 550–562.
   Web of Science ®Google Scholar
• Pictet, F. J., & Humbert, A. (1857). Description d'une émyde nouvelle (Emys etalloni) du terrain Jurassique supérieur des environs de St. Claude (Vol. 1). Kessmann.
   Google Scholar
• Rabi, M., Sukhanov, V. B., Egorova, V. N., Danilov, I. G., & Joyce, W. G. (2014). Osteology, relationships, and ecology of Annemys (Testudines, Eucryptodira) from the Late Jurassic of Shar Teg, Mongolia, and phylogenetic definitions for Xinjiangchelyidae, Sinemydidae, and Macrobaenidae. Journal of Vertebrate Paleontology, 34(2), 327–352.
   Web of Science ®Google Scholar
• Rabi, M., Zhou, C. F., Wings, O., Ge, S., & Joyce, W. G. (2013). A new xinjiangchelyid turtle from the Middle Jurassic of Xinjiang, China and the evolution of the basipterygoid process in Mesozoic turtles. BMC Evolutionary Biology, 13(1), 1–29.
   PubMedGoogle Scholar
• Rollot, Y., Evers, S. W., & Joyce, W. G. (2021a). A redescription of the Late Jurassic (Tithonian) turtle Uluops uluops and a new phylogenetic hypothesis of Paracryptodira. Swiss Journal of Palaeontology, 140(1), 1–30.
   Web of Science ®Google Scholar
• Rollot, Y., Evers, S. W., & Joyce, W. G. (2021b). A review of the carotid artery and facial nerve canal systems in extant turtles. PeerJ, 8, e10475.
   PubMedGoogle Scholar
• Romer, A. S. (1956). The Osteology of the Reptiles. University Chicago Press.
   Google Scholar
• Rougier, G. W., de la Fuente, M. S., & Arcucci, A. B. (1995). Late Triassic turtles from South America. Science, 268(5212), 855–858.
   PubMed Web of Science ®Google Scholar
• Scheyer, T. M., & Sander, P. M. (2007). Shell bone histology indicates terrestrial palaeoecology of basal turtles. Proceedings of the Royal Society B: Biological Sciences, 274(1620), 1885–1893.
   PubMed Web of Science ®Google Scholar
• Scheyer, T. M., Danilov, I. G., Sukhanov, V. B., & Syromyatnikova, E. V. (2014). The shell bone histology of fossil and extant marine turtles revisited. Biological Journal of the Linnean Society, 112(4), 701–718.
   Web of Science ®Google Scholar
• Skutschas, P. P., Markova, V. D., Kolchanov, V. V., Averianov, A. O., Martin, T., Schellhorn, R., Kolosov, P. N., Grigoriev, D. V., Vitenko, D. D., Obraztsova, E. M., & Danilov, I. G. (2020). Basal turtle material from the Lower Cretaceous of Yakutia (Russia) filling the gap in the Asian record. Cretaceous Research, 106(104186).
   Web of Science ®Google Scholar
• Sterli, J. (2008). A new, nearly complete stem turtle from the Jurassic of South America with implications for turtle evolution. Biology Letters, 4(3), 286–289.
   PubMed Web of Science ®Google Scholar
• Sterli, J. (2010). Phylogenetic relationships among extinct and extant turtles: the position of Pleurodira and the effects of the fossils on rooting crown-group turtles. Contributions to Zoology, 79(3), 93–106.
   Web of Science ®Google Scholar
• Sterli, J., & de la Fuente, M. S. (2010). Anatomy of Condorchelys antiqua Sterli, 2008, and the origin of the modern jaw closure mechanism in turtles. Journal of Vertebrate Paleontology, 30(2), 351–366.
   Web of Science ®Google Scholar
• Sterli, J., & Joyce, W. G. (2007). The cranial anatomy of the Early Jurassic turtle Kayentachelys aprix. Acta Palaeontologica Polonica, 52(4), 675–694.
   Web of Science ®Google Scholar
• Sterli, J., de la Fuente, M., & Rougier, G. W. (2007). Anatomy and relationships of Palaeochersis talampayensis, a Late Triassic turtle from Argentina. Palaeontographica Abteilung A, 281, 1–61.
   Google Scholar
• Sterli, J., de la Fuente, M. S., & Rougier, G. W. (2018). New remains of Condorchelys antiqua (Testudinata) from the Early–Middle Jurassic of Patagonia: anatomy, phylogeny, and paedomorphosis in the early evolution of turtles. Journal of Vertebrate Paleontology, 38(4), 1–17.
   Web of Science ®Google Scholar
• Sterli, J., Müller, J., Anquetin, J., & Hilger, A. (2010). The parabasisphenoid complex in Mesozoic turtles and the evolution of the testudinate basicranium. Canadian Journal of Earth Sciences, 47(10), 1337–1346.
   Web of Science ®Google Scholar
• Sukhanov, V. B. (1964). Subclass Testudinata. In Yu. A. Orlov (Ed.), Osnovy Paleontologii. Zemnovodnye, Presmykayushchiesya i Ptitsy [Fundamentals of Paleontology. Amphibians, Reptiles, and Birds] (pp. 354–438). Nauka, Moscow. [in Russian].
   Google Scholar
• Sukhanov, V. B. (2000). Mesozoic turtles of Middle and Central Asia. In M. J. Benton, M. A. Shishkin, D. M. Unwin, & E. N. Kurochkin (Eds.), The Age of Dinosaurs in Russia and Mongolia (pp. 309–367). Cambridge University Press.
   Google Scholar
• Sukhanov, V. B. (2001). An archaic turtle from the Middle Jurassic of Moskovskaya Region and its position in basal radiation of the order Testudines. In Voprosy gerpetologii. Materialy I s’’ezda Gerpetologicheskogo ob-va im. A.M. Nikol’skogo [I Congress of the A.M. Nikolsky Herpetological Society on the Questions of Herpetology] (pp. 282–284). Pushchino–Moscow. [in Russian]
   Google Scholar
• Sukhanov, V. B. (2003). Heckerochelys romani gen. et sp. nov., an archaic turtle, from the Middle Jurassic of the Moscow Region and its place in the basal radiation of Testudines. In Symposium on Turtle Origins, Evolution and Systematics. Program and Abstracts (pp. 29–30). St. Petersburg, Russia.
   Google Scholar
• Sukhanov, V. B. (2006). An archaic turtle, Heckerochelys romani gen. et sp. nov., from the Middle Jurassic of Moscow region, Russia. Fossil Turtle Research, 1 (Russian Journal of Herpetology, 13, Suppl.), 112–118.
   Google Scholar
• Sukhanov, V. B., & Narmandakh, P. (2006). New taxa of Mesozoic turtles from Mongolia. Fossil turtle research, 1, 119–127.
   Google Scholar
• Suzuki, S., & Chinzorig, T. (2010). A catalog of Mongolochelys collected by the HMNS-MPC Joint Paleontological Expedition. Hayashibara Museum of Natural Sciences Research Bulletin, 3, 119–131.
   Google Scholar
• Tatarinov, L. P. (1959). Features of the sound-conducting apparatus of fossil and modern turtles. Paleontologicheskii Zhurnal, 3, 112–116. [in Russian]
   Google Scholar
• Warraich, N., Wyneken, J., & Blume, N. (2020). Feeding behavior and visual field differences in loggerhead and leatherback sea turtles may explain differences in longline fisheries interactions. Endangered Species Research, 41, 67–77.
   Web of Science ®Google Scholar
• Werneburg, I. (2012). Temporal bone arrangements in turtles: an overview. Journal of Experimental Zoology Part B: Molecular and Developmental Evolution, 318(4), 235–249.
   PubMedGoogle Scholar
• Werneburg, I., & Maier, W. (2019). Diverging development of akinetic skulls in cryptodire and pleurodire turtles: an ontogenetic and phylogenetic study. Vertebrate Zoology, 69(2), 113–143.
   Web of Science ®Google Scholar
• Willis, K. L., Christensen-Dalsgaard, J., Ketten, D. R., & Carr, C. E. (2013). Middle ear cavity morphology is consistent with an aquatic origin for testudines. PLoS ONE, 8(1), e54086.
   PubMed Web of Science ®Google Scholar
• Wings, O., Rabi, M., Schneider, J. W., Schwermann, L., Sun, G., Zhou, C. F., & Joyce, W. G. (2012). An enormous Jurassic turtle bone bed from the Turpan Basin of Xinjiang, China. Naturwissenschaften, 99(11), 925–935.
   PubMedGoogle Scholar