A new Tithonian ophthalmosaurid ichthyosaur from Coahuila in northeastern Mexico

References

• Alvarado-Ortega, J., Barrientos-Lara, J.I., Espinosa-Arrubarrena, L. & Melgarejo-Damián, M.P., 2014. Late Jurassic marine vertebrates from Tlaxiaco, Oaxaca State, southern Mexico. Palaeontologia Electronica 17, 1–25. palaeo-electronica.org/content/2014/773-tlaxiaco-jurassic-vertebrates.
   Web of Science ®Google Scholar
• Arkhangelsky, M.S., 2001. On a new ichthyosaur of the genus Otschevia from the Volgian stage of the Volga region near Ulyanovsk. Paleontological Journal 35, 629–634.
   Google Scholar
• Bardet, N., Falconnet, J., Fischer, V., Houssaye, A., Jouve, S., Pereda Suberbiola, X., Pérez-García, A., Rage, J.-C. & Vincent, P., 2014. Mesozoic marine reptile palaeobiogeography in response to drifting plates. Gondwana Research 26, 869–887.
   Web of Science ®Google Scholar
• Bardet, N. & FernÁndez, M., 2000. A new Ichthyosaur from the Upper Jurassic Lithographic Limestones of Bavaria. Journal of Paleontology 74, 503–511.
   Web of Science ®Google Scholar
• Barrientos-Lara, J.I. & Alvarado-Ortega, J., 2020. Acuetzpalin carranzai gen et sp. nov. A new ophthalmosauridae (Ichthyosauria) from the Upper Jurassic of Durango, North Mexico. Journal of South American Earth Sciences 98, 102456.
   Web of Science ®Google Scholar
• Barrientos-Lara, J.I., Alvarado-Ortega, J. & FernÁndez, M.S., 2018. The marine crocodile Maledictosuchus (Thalattosuchia, Metriorhynchidae) from the Kimmeridgian Deposits of Tlaxiaco, Oaxaca, Southern Mexico. Journal of Vertebrate Paleontology 38, 1–14.
   Web of Science ®Google Scholar
• Barrientos-Lara, J.I., FernÁndez, M.S., &. & Alvarado-Ortega, J., 2015. Kimmeridgian pliosaurids (Sauropterygia, Plesiosauria) from Tlaxiaco, Oaxaca, southern Mexico. Revista Mexicana de Ciencias Geológicas 32, 293–304.
   Web of Science ®Google Scholar
• Barrientos-Lara, J.I., Herrera, Y., FernÁndez, M.S. & Alvarado-Ortega, J., 2017. Occurrence of Torvoneustes (Crocodylomorpha, Metriorhynchidae) in marine Jurassic deposits of Oaxaca, Mexico. Revista Brasileira de Paleontologia 19, 415–424.
   Web of Science ®Google Scholar
• Baur, G., 1887. On the morphology and origin of the Ichthyopterygia. American Naturalist 21, 837–840.
   Google Scholar
• Boulenger, G.A., 1904. Exhibition of, and remarks upon, a paddle of a new species of ichthyosaur. Proceedings of the Zoological Society of London 1904, 424–426.
   Google Scholar
• Bremer, K., 1994. Branch support and tree stability. Cladistics 10, 295–304.
   Web of Science ®Google Scholar
• Broili, F., 1907. Ein neuer Ichthyosaurus aus der norddeutschen Kreide. Palaeontographica 54, 139–162.
   Google Scholar
• Buchy, M.C., 2007. Mesozoic Marine Reptiles from North-East Mexico: Description, Systematics, Assemblages and Palaeobiogeography. PhD thesis, University of Karlsruhe, Germany, 98 pp. (unpublished)
   Google Scholar
• Buchy, M.C., 2010. First record of Ophthalmosaurus (Reptilia: Ichthyosauria) from the Tithonian (Upper Jurassic) of Mexico. Journal of Paleontology 84, 149–155.
   Web of Science ®Google Scholar
• Buchy, M.C., Frey, E., Stinnesbeck, W. & López-Oliva, J.G., 2006. An annotated catalogue of the Upper Jurassic (Kimmeridgian and Tithonian) marine reptiles in the collections of the Universidad Autónoma de Nuevo León, Facultad de Ciencias de la Tierra, Linares, Mexico. Oryctos 6, 1–18.
   Google Scholar
• Buchy, M.C. & López-Oliva, J.G., 2009. Occurrence of a second ichthyosaur genus (Reptilia; Ichthyosauria) in the Late Jurassic Gulf of Mexico. Boletín de la Sociedad Geológica Mexicana 61, 233–238.
   Google Scholar
• Campos, L., FernÁndez, M.S. & Herrera, Y., 2019. A new ichthyosaur from the Late Jurassic of north-west Patagonia (Argentina) and its significance for the evolution of the narial complex of the ophthalmosaurids. Zoological Journal of the Linnean Society 188, 180–201.
   Web of Science ®Google Scholar
• Conybeare, W.D., 1822. Additional notices on the fossil genera Ichthyosaurus and Plesiosaurus. Transactions of the Geological Society of London s2-1, 103–123.
   Google Scholar
• De Blainville, H.D., 1835. Description de quelques espèces de reptiles de la Californie, preécédé de l’analyse d’un système général d’erpé tologie et d’amphibiologie. Nouvelles Annales du Muséum D’Histoire Naturelle de Paris 4, 233–296.
   Google Scholar
• De La Beche, H.T. & Conybeare, W.D., 1821. Notice of the discovery of a new Fossil Animal, forming a link between the Ichthyosaurus and Crocodile, together with general remarks on the Osteology of the Ichthyosaurus. Transactions of the Geological Society of London S1-5, 559–594.
   Google Scholar
• Delsett, L.L., Roberts, A.J., Druckenmiller, P.S. & Hurum, J.H., 2017. A new ophthalmosaurid (Ichthyosauria) from Svalbard, Norway, and evolution of the ichthyopterygian pelvic girdle. PLoS ONE 12, e0169971.
   PubMed Web of Science ®Google Scholar
• Delsett, L.L., Roberts, A.J., Druckenmiller, P.S. & Hurum, J.H., 2019. Osteology and phylogeny of Late Jurassic ichthyosaurs from the Slottsmøya Member Lagerstätte (Spitsbergen, Svalbard). Acta Palaeontologica Polonica 64, 717–743.
   Web of Science ®Google Scholar
• Druckenmiller, P.S., Hurum, J.H., Knutsen, E.M. & Nakrem, H.A., 2012. Two new ophthalmosaurids (Reptilia: Ichthyosauria) from the Agardhfjellet Formation (Upper Jurassic: Volgian/Tithonian), Svalbard, Norway. Norwegian Journal of Geology 92, 311–339.
   Web of Science ®Google Scholar
• Druckenmiller, P.S. & Maxwell, E.E., 2010. A new Lower Cretaceous (lower Albian) ichthyosaur from the Clearwater Formation, Alberta, Canada. Canadian Journal of Earth Sciences 47, 1037–1053.
   Web of Science ®Google Scholar
• Efimov, V.M., 1999. A new family of ichthyosaurs, the Undorosauridae fam. nov. from the Volgian stage of the European part of Russia. Paleontological Journal 33, 174–181.
   Google Scholar
• Felsenstein, J., 1985. Confidence limits on phylogenies: an approach using the bootstrap. Evolution 39, 783–791.
   PubMed Web of Science ®Google Scholar
• FernÁndez, M.S., 1994. A new long-snouted ichthyosaur from the Early Bajocian of Neuquén Basin (Argentina). Ameghiniana 31, 291–297.
   Google Scholar
• FernÁndez, M.S., 1997. A new ichthyosaur from the Tithonian (Late Jurassic) of the Neuquén Basin (Argentina). Journal of Paleontology 71, 479–484.
   Web of Science ®Google Scholar
• FernÁndez, M.S., 1999. A new ichthyosaur from the Los Molles Formation (Early Bajocian), Neuquen Basin, Argentina. Journal of Paleontology 73, 677–681.
   Web of Science ®Google Scholar
• FernÁndez, M.S. & Agirre-Urreta, M.B., 2005. Revision of Platypterygius hauthali von Huene, 1927 (Ichthyosauria: Ophthalmosauridae) from the Early Cretaceous of Patagonia, Argentina. Journal of Vertebrate Paleontology 25, 583–587.
   Web of Science ®Google Scholar
• FernÁndez, M. & Iturralde-Vinent, M., 2000. An Oxfordian ichthyosauria (Reptilia) from Viñales, western Cuba: paleobiogeographic significance. Journal of Vertebrate Paleontology 20, 191–193.
   Web of Science ®Google Scholar
• FernÁndez, M.S. & Talevi, M., 2014. Ophthalmosaurian (Ichthyosauria) records from the Aalenian-Bajocian of Patagonia (Argentina): an overview. Geological Magazine 151, 49–11.
   Web of Science ®Google Scholar
• Fischer, V., 2012. New data on the ichthyosaur Platypterygius hercynicus and its implications for the validity of the genus. Acta Palaeontologica Polonica 57, 123–134.
   Web of Science ®Google Scholar
• Fischer, V., Appleby, R.M., Naish, D., Liston, J., Riding, J.B., Brindley, S. & Godefroit, P., 2013. A basal thunnosaurian from Iraq reveals disparate phylogenetic origins for Cretaceous ichthyosaurs. Biology Letters 9, 20130021.
   PubMed Web of Science ®Google Scholar
• Fischer, V., Bardet, N., Guiomar, M. & Godefroit, P., 2014a. High diversity in Cretaceous ichthyosaurus from Europe prior to their extinction. PLoS One 9, e84709.
   PubMed Web of Science ®Google Scholar
• Fischer, V., Arkhangelsky, M.S., Uspensky, G.N., Sten-Shin, I.M. & Godefroit, P., 2014b. A new Lower Cretaceous ichthyosaur from Russia reveals skull shape conservatism within Ophthalmosaurinae. Geological Magazine 151, 60–70.
   Web of Science ®Google Scholar
• Fischer, V., Arkhangelsky, M.S., Naish, D., Stenshin, I.M., Uspensky, G.N. & Godefroit, P., 2014c. Simbirskiasaurus and Pervushovisaurus reassessed: implications for the taxonomy and cranial osteology of Cretaceous platypterygiine ichthyosaurs. Zoological Journal of the Linnean Society 171, 822–841.
   Web of Science ®Google Scholar
• Fischer, V., Bardet, N., Benson, R.B.J., Arkhangelsky, M.S. & Friedman, M., 2016. Extinction of fish-shaped marine reptiles associated with reduced evolutionary rates and global environmental volatility. Nature Communications 7, 10825.
   Web of Science ®Google Scholar
• Fischer, V., Maisch, M.W., Naish, D., Kosma, R., Liston, J., Joger, U., KrÜger, F.J., Fritz, J., Pardo Pérez, J., Tainsh, J., &. & Appleby, R., 2012. New Ophthalmosaurid ichthyosaurs from the European Lower Cretaceous demonstrate extensive ichthyosaur survival across the Jurassic–Cretaceous boundary. PLoS One 7, e29234.
   PubMed Web of Science ®Google Scholar
• Fischer, V., Masure, E., Arkhangelsky, M.S. & Godefroit, P., 2011. A new Barremian (Early Cretaceous) ichthyosaur from western Russia. Journal of Vertebrate Paleontology 31, 1010–1025.
   Web of Science ®Google Scholar
• Frey, E. & Strinnesbeck, W., 2014. Mexican Ichthyosaurs. In Dinosaurs and Other Reptiles from the Mesozoic. Rivera-Silva, H.E. & Carpenter, K. Frey, E., eds, Indiana University Press, Indianapolis, 99–109.
   Google Scholar
• Gasparini, Z. & Iturralde-Vinent, M., 2006. The Cuban Oxfordian herpetofauna in the Caribbean Seaway. Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen 240, 343–371.
   Web of Science ®Google Scholar
• Goloboff, F. & Catalano, S., 2016. TNT, version 1.5, with a full implementation of phylogenetic morphometrics. Cladistics 32, 221–238.
   PubMed Web of Science ®Google Scholar
• Huene, F.V., 1922. Die Ichthyosaurier des Lias und ihre Zusammenhange. Monographien zur Geologic and Palaeontologie 1, 1–114.
   Google Scholar
• Huene, F.V., 1927. Beitrag zur Kenntnis mariner mesozoisher Wirbeltiere in Argentinien. Centralblatt für Mineralogie, Geologie und Paläontologie B 22–29.
   Google Scholar
• Hulke, J.W., 1871. Note on an Ichthyosaurus (I. enthekiodon) from Kimmeridge Bay, Dorset. Quarterly Journal of the Geological Society of Society 27, 440–441.
   Google Scholar
• Kear, B.P., 2003. Cretaceous marine reptiles of Australia: a review of taxonomy and distribution. Cretaceous Research 24, 277–303.
   Web of Science ®Google Scholar
• Kear, B.P., 2005. Cranial morphology of Platypterygius longmani Wade, 1990 (Reptilia: Ichthyosauria) from the Lower Cretaceous of Australia. Zoological Journal of the Linnean Society 145, 583–622.
   Web of Science ®Google Scholar
• Kear, B.P. & Zammit, M., 2014. In utero foetal remains of the Cretaceous ichthyosaurian Platypterygius: ontogenetic implications for character state efficacy. Geological Magazine 151, 71–86.
   Web of Science ®Google Scholar
• Kirton, A.M., 1983. A Review of British Upper Jurassic Ichthyosaurs. PhD thesis, University of Newcastle-upon-Tyne, 239 pp. (Unpublished)
   Google Scholar
• Kolb, C. & Sander, P.M., 2009. Redescription of the ichthyosaur Platypterygius hercynicus (Kuhn 1946) from the Lower Cretaceous of Salzgitter (Lower Saxony, Germany). Palaeontographica Abteilung A 288, 151–192.
   Google Scholar
• Kuhn, O., 1946. Ein Skelett von Ichthyosaurus (Platypterygius) hercynicus n. sp. aus dem Aptium von Gitter. Berichte Der Naturforschenden Gesellschaft Bamberg 29, 69–82.
   Google Scholar
• Lomax, D.R. & Massare, J.A., 2017. Two new species of ichthyosaurus from the lowermost Jurassic (Hettangian) of somerset, England. Papers in Palaeontology 3, 1–20.
   Web of Science ®Google Scholar
• Maddison, W.P. & Maddison, D.R., 2018. Mesquite: a modular system for evolutionary analysis. Version 3 31. http://mesquiteproject.org.
   Google Scholar
• Maxwell, E.E., 2010. Generic reassignment of an ichthyosaur from the Queen Elizabeth Islands, Northwest Territories. Journal of Vertebrate Paleontology 30, 403–415.
   Web of Science ®Google Scholar
• Maxwell, E.E. & Caldwell, M.W., 2006. A new genus of ichthyosaur from the Lower Cretaceous of Western Canada. Palaeontology 49, 1043–1052.
   Web of Science ®Google Scholar
• Maxwell, E.E. & Cortés, D., 2020. A revision of the Early Jurassic ichthyosaur Hauffiopteryx (Reptilia: Ichthyosauria), and description of a new species from southwestern Germany. Palaeontologia Electronica 23, a31.
   Web of Science ®Google Scholar
• Maxwell, E.E., Cortés, D., Patarroyo, P. & Parra-Ruge, M.L., 2019. A new specimen of Platypterygius sachicarum (Reptilia, Ichthyosauria) from the Early Cretaceous of Colombia and its phylogenetic implications. Journal of Vertebrate Paleontology 39, e1577875.
   Web of Science ®Google Scholar
• Maxwell, E.E., Dick, D., Padilla, S. & Parra M.L., 2016. A new opthalmosaurid ichthyosaur from the Early Cretaceous of Colombia. Papers in Palaeontology 2, 59–12.
   Web of Science ®Google Scholar
• Maxwell, E.E., FernÁndez, M.S. & Schoch, R.R., 2012. First diagnostic marine reptile remains from the Aalenian (Middle Jurassic): a new ichthyosaur from south western Germany. PLoS One 7, e41692.
   PubMed Web of Science ®Google Scholar
• Maxwell, E.E. & Kear, B.P., 2010. Postcranial anatomy of Platypterygius americanus (Reptilia: Ichthyosauria) from the Cretaceous of Wyoming. Journal of Vertebrate Paleontology 30, 1059–1068.
   Web of Science ®Google Scholar
• MccOy, F., 1867. On the occurrence of Ichthyosaurus and Plesiosaurus in Australia. Annals and Magazine of Natural History 19, 355–356.
   Google Scholar
• McGowan, C., 1972. The systematics of Cretaceous ichthyosaurs with particular reference to the material from North America. Contributions to Geology, University of Wyoming 11, 9–29.
   Google Scholar
• McGowan, C., 1974. A revision of the longipinnate icthyosaurus of the Lower Jurassic of England, with descriptions of two new species (Reptilia: Ichthyosauria). Life Sciences Contributions, Royal Ontario Museum 97, 1–37.
   Google Scholar
• McGowan, C., 1976. The description and phenetic relationships of a new ichthyosaur genus from the Upper Jurassic of England. Canadian Journal of Earth Sciences 13, 668–683.
   Web of Science ®Google Scholar
• McGowan, C. & Motani, R., 2003. Ichthyopterygia. Handbook of Paleoherpetology Part 8. Verlag Dr. Friedrich Pfeil, Münich, 175 pp.
   Google Scholar
• Moon, B.C., 2017. A new phylogeny of ichthyosaurs (Reptilia: Diapsida). Journal of Systematic Paleontology 17, 129–155.
   Google Scholar
• Moon, B.C. & Kirton, A.M., 2016. Ichthyosaurs of the British Middle and Upper Jurassic Part 1, Ophthalmosaurus. Monographs of the Palaeontographical Society 170, 1–84.
   Google Scholar
• Moon, B.C. & Kirton, A.M., 2018. Ichthyosaurs of the British Middle and Upper Jurassic Part 2. Brachypterygius, nannopterygius, macropterygius and taxa invalida. Monographs of the Palaeontographical Society 172, 85–197.
   Google Scholar
• Motani, R., 1999. On the evolution and homologies of ichthyosaurian forefins. Journal of Vertebrate Paleontology 19, 28–41.
   Web of Science ®Google Scholar
• Nace, R.L., 1939. A new ichthyosaur from the Upper Cretaceous Mowry Formation of Wyoming. American Journal of Science 237, 673–686.
   Google Scholar
• Ochev, V.G. & Efimov, V.M., 1985. A new genus of Ichthyosaur from the Ul’yanovsk area of the Povolzh’ye Region. Paleontological Journal 4, 87–91.
   Google Scholar
• Paparella, I., Maxwell, E.E., Cipriani, A., Roncacè, S. & Caldwell, M.W., 2017. The first ophthalmosaurid ichthyosaur from the Upper Jurassic of the Umbrian–Marchean Apennines (Marche, Central Italy). Geological Magazine 154, 837–822.
   Web of Science ®Google Scholar
• PÁramo, M.E., 1997. Platypterygius sachicarum (Reptilia, Ichthyosauria) nueva especie del Cretácico de Colombia. Revista Ingeominas 6, 1–12.
   Google Scholar
• Prasad, G.V.R., Pandey, D.K., Alberti, M., FÜrsich, F.T., Thakkar, M.G. & Chauhan, G.D., 2017. Discovery of the first ichthyosaur from the Jurassic of India: implications for Gondwanan palaeobiogeography. PLoS ONE 12, e0185851.
   PubMed Web of Science ®Google Scholar
• Quenstedt, F.A., 1852. Handbuch der Petrefaktenkunde. H. Laupp, Tübingen IV, 792 pp.
   Google Scholar
• Roberts, A.J., Druckenmiller, P.S., Saetre, G.-P. & Hurum, J.H., 2014. A new Upper Jurassic ophthalmosaurid ichthyosaur from the Slottsmøya Member, Agardhfjellet Formation or Central Spitsbergen. PLoS One 9, e103152.
   PubMed Web of Science ®Google Scholar
• Russell, D.A., 1993. Jurassic marine reptiles from Cape Grassy, Melville Island, Arctic Canada. The Geology of Melville Island, Arctic Canada. Geological Survey of Canada Bulletin 450, 195–201.
   Google Scholar
• Seeley, H.G., 1874. On the pectoral arch and forelimb of Ophthalmosaurus, a new ichthyosaurian genus from the Oxford Clay. Quarterly Journal of the Geological Society 30, 696–707.
   Google Scholar
• Servicio GeolÓgico Mexicano. 2008. Carta Geológica-Minera, Monterrey G14-7, Nuevo León y Coahuila 1: 250000. 2da ed. Secretaria de Economia, México.
   Google Scholar
• Smith, P.L., 1983. The Pliensbachian ammonite Dayiceras dayiceroides and Early Jurassic paleogeography. Canadian Journal of Earth Sciences 20, 86–91.
   Web of Science ®Google Scholar
• Tyborowski, D., 2016. A new Ophthalmosaurid Ichthyosaur from the Late Jurassic of Owadów-Brzezinki Quarry, Poland. Acta Palaeontologica Polonica 61, 791–803.
   Web of Science ®Google Scholar
• Wagner, A., 1853. Die Characteristic einer neuen Art von Ichthyosaurus aus den lithographischen Schiefern und eines Zahnes von Polyptychodon aus dem Gründsandsteine von Kelheim. Bulletin der Königlische Akademie der Wissenschaft, Gelehrte Anzeigen 3, 25–35.
   Google Scholar
• Zammit, M., 2010. A review of Australasian ichthyosaurs. Alcheringa: An Australasian Journal of Palaeontology 34, 281–292.
   Google Scholar
• Zammit, M., Norri, S.R.M. & Kear, B.P., 2010. The Australian Cretaceous ichthyosaur Platypterygius australis: a description and review of postcranial remains. Journal of Vertebrate Paleontology 30, 1726–1735.
   Web of Science ®Google Scholar
• Zverkov, N.G., Arkhangelsky, M.S., Pardo-Pérez, J.M. & Beznosov P, A., 2015. On the Upper Jurassic Ichthyosaur remains from the Russian North. Proceedings of the Zoological Institute RAS 319, 81–97.
   Google Scholar
• Zverkov, N.G. & Efimov, V.M., 2019. Revision of Undorosaurus, a mysterious Late Jurassic ichthyosaur of the Boreal Realm. Journal of Systematic Palaeontology 17, 1183–1213.
   Web of Science ®Google Scholar
• Zverkov, N.G. & Grigoriev, D.V., 2020. An unrevealed lineage of platypterygiines (Ichthyosauria) with peculiar forefin structure and semiglobal distribution in the mid-Cretaceous (Albian–Cenomanian). Cretaceous Research 115, 104550–104517.
   Web of Science ®Google Scholar
• Zverkov, N.G. & Jacobs, M.L., 2021. Revision of Nannopterygius (Ichthyosauria: Ophthalmosauridae): reappraisal of the ‘inaccessible’ holotype resolves a taxonomic tangle and reveals an obscure ophthalmosaurid lineage with a wide distribution. Zoological Journal of the Linnean Society 191, 228–274.
   Web of Science ®Google Scholar
• Zverkov, N.G. & Prilepskaya, N.E., 2019. A prevalence of Arthropterygius (Ichthyosauria: Ophthalmosauridae) in the Late Jurassic-earliest Cretaceous of the Boreal Realm. PeerJ 7, e6799.
   PubMed Web of Science ®Google Scholar