- Bennett , S.C. 1991 . Morphology of the late Cretaceous pterosaur Pteranodon and systematics of the Pterodactyloidea , Lawrence, Kansas : University of Kansas . Unpublished Ph.D. dissertation
- Bennett , S.C. 1996 . The phylogenetic position of the Pterosauria within the Archosauromorpha . Zoological Journal of the Linnean Society , 118 : 261 – 308 .
- Bennett , S.C. 1997 . The arboreal leaping theory of the origin of pterosaur flight . Historical Biology , 29 : 373 – 386 .
- Bennett , S.C. 2000a . Pterosaur flight: the role of actinofibrils in wing function . Historical Biology , 14 : 255 – 284 .
- Bennett , S.C. 2000b . The osteology and functional morphology of the late Cretaceous pterosaur Pteranodon. Part I. General description of osteology. Part II. Size and functional morphology . Palaeontographica A (Stuttgart) , 260 : 1 – 153 .
- Benton , M.J. 1982 . The Diapsida: revolution in reptile relationships . Nature , 296 : 306 – 307 .
- Benton , M.J. 1984 . The relationships and early evolution of the Diapsida . Symposia of the Zoological Society of London , 52 : 575 – 596 .
- Benton , M.J. 1985 . Classification and phytogeny of the diapsid reptiles . Zoological Journal of the Linnean Society , 84 : 97 – 164 .
- Benton , M.J. 1999 . Scleromochlus taylori and the origin of dinosaurs and pterosaurs . Philosophical Transactions of the Royal Society, London, B , 354 : 1423 – 1446 .
- Boonstra , L.D. 1967 . An early stage in the evolution of the mammalian quadrupedal walking gait . Annals of the South African Museum , 50 (3) : 27 – 42 .
- Brown , R.E. , Baumel , J.J. and Klemm , R.D. 1995 . Mechanics of the avian propatagium: flexion-extension mechanism of the avian wing . Journal of Morphology , 225 : 91 – 105 .
- Buckland , W. 1835 . On the discovery of a new species of pterodactyl in the Lias at Lyme Regis . Transactions of the Geological Society, London, series 2 , 3 : 217 – 222 .
- Clark , J. , Hopson , J. , Hernandez , R. , Fastovsky , D. and Montellano , M. 1998 . Foot posture in a primitive pterosaur . Nature , 391 : 886 – 889 .
- Clarke , J.A. , Gauthier , J.A. , Norell , M.A. and Ji , Q. 2001 . The origin and significance of a propatagium in flying dinosaurs . Journal of Vertebrate Paleontology , 21 (Supplement to No. 3) : 41A
- Cox , M.J. , van Dijk , P.P. , Nabhitabhata , J. and Thirakhupt , K. 1998 . A Photographic Guide to Snakes and Other Reptiles of Peninsular Malaysia, Singapore and Thailand , Sanibel Island, Florida : Ralph Curtis Publishing .
- Dalla Vecchia , F.M. 1995 . A new pterosaur (Reptilia, Pterosauria) from the Norian (Late Triassic) of Friuli (northeastern Italy). Preliminary note . Gortania, Atti del Museo Friulano di Storia Naturale , 16 : 59 – 66 .
- Dial , K. 2001 . On the origin and ontogeny of avian flight: wing-assisted incline running . Journal of Vertebrate Paleontology , 21 (Supplement to No. 3) : 45A
- Ebe , L K. 1996 . On the origin of flight in Archaeopteryx and in pterosaurs . Neues Jahrbuch für Geologie and Paläontologie, Abhandlung , 202 : 269 – 285 .
- Ellenberger , P. and de Villalta , J.F. 1974 . Sur la presence d'un ancêtre probable des oiseaux dans le Muschelkalk supérieure de Catalogue (Espagne). Note préliminaire . Acta Geologien Hispanica , 9 : 162 – 168 .
- Frey , E. and Reiss , J. 1981 . A new reconstruction of the pterosaur wing . Neues Jahrbuch far Geologie and Paläontologie, Abhandlung , 161 : 1 – 27 .
- Frey , E. and Martill , D. 1994 . A new pterosaur from the Crato formation (Lower Cretaceous Aptian) of Brazil . Neues Jahrbuch für Geologie und Paläontologie, Abhandlung , 194 : 379 – 412 .
- Gans , C. , Darevksi , I. and Tatarinov , L.P. 1987 . Sharovipteryx, a reptilian glider? . Paleobiology , 13 : 415 – 426 .
- Gatesy , S.M. and Dial , K.P. 1996 . Locomotor modules and the evolution of avian flight . Evolution , 50 : 331 – 340 .
- Gauthier , J.A. 1984 . A Cladistic Analysis of the Higher Systematic Categories of the Diapsida Unpublished , Berkeley, California : University of California . PKD. dissertation
- Gauthier , J. A. 1986 . “ Saurischian monophyly and the origin of birds ” . In The Origin of Birds and the Evolution of Flight , Edited by: Padian , K. Vol. 8 , 1 – 55 . San Francisco : Memoirs of the California Academy of Sciences .
- Haubold , H. and Buffetaut , E. 1987 . Une nouvelle interprétation de Longisquama insignis, reptile énigmatique du Trias supériuer d'Asie central . Comptes rendus de l'Académie des Sciences de Paris , 305 : 65 – 70 .
- Huene , F. von . 1914 . Beiträge zur geschiente der archosaurier . Geologische and palaeontokgische Abhandlungen N.F. , 13 : 1 – 53 .
- Huene , F. von . 1921 . Neue Pseudosuchier und Coelurosaurier aus dem Württembergischen Keuper . Acta Zoologien , 2 : 329 – 403 .
- Jenkins , FA. 1971 . The posteranial skeleton of African cynodonts . Bulletin of the Peabody Museum of Natural History, Yale University , 36 : 1 – 216 .
- Jenkins , FA. , Shubin , N.H. , Gatesy , S.M. and Padian , K. 2001 . A diminutive pterosaur (Pterosauria: Eudimorphodontidae) from the Greenlandic Triassic Bulletin of the Museum of Comparative Zoology . Harvard University , 155 (9) : 487 – 506 .
- Jones , T.D. , Ruben , J.A. , Martin , L.D. , Kurochkin , E.N. , Feduccia , A. , Maderson , P.F.A. , Hillenius , W.J. , Geist , N.R. and Alifanov , V. 2000 . Non-avian feathers in a Late Triassic archosaur . Science , 288 : 2202 – 2204 .
- Lockley , MG. , Logue , T.J. , MorataUa , J.J. , Hunt , A.P. , Schultz , RJ. and Robinson , J.W. 1995 . The fossil trackway Pteraichnus is pterosaurian, not crocodilian; implications for the global distribution of pterosaur tracks . Ichnos , 4 : 7 – 20 .
- Lu , J. and Wang , X. 2001 . Soft tissue in an Early Cretaceous pterosaur from Liaoning Province, China . Journal of Vertebrate Paleontology , 21 (Supplement to No. 3) : 74a
- Mazin , J.-M. , Hantzpergue , P. , Lafaurie , G. and Vignaud , P. 1995 . Des pistes de ptérosaures dans le Tithonien de Crayssac (Quercy, France) . Comptes rendus de l'Académie des Sciences de Paris , 321 : 417 – 424 .
- Mazin , J.-M. , Billon-Buyat , J.-P , Hantzpergue , P. and LaFaurie , G. 2001 . “ Could they be the first rhamphorhynchoid tracks? Yes! ” . In Two Hundred Years of Pterosaurs , Edited by: Sabatier , P. 1 – 11 . 64 – 65 . Strata Série : Laboratoire de Géologie sédimentaire et Paléolontologie, Université de Toulouse .
- Owen , R. 1870 . A Monograph on the Fossil Reptilia of the Liassic Formations. III. Part U: Order Pterosauria , 41 – 81 . London : Palaeontolographical Society .
- Padian , K. 1979 . The wings of pterosaurs: a new look . Discovery , 14 : 21 – 29 .
- Padian , K. 1980 . Studies of the Structure, Evolution and Flight of Pterosaurs (Reptilia, Pterosauria) , New Haven, Connecticut : Yale University . Unpublished Ph.D. dissertation
- Padian , K. 1983 . A functional analysis of flying and walking in pterosaurs . Paleobiology , 9 : 218 – 239 .
- Padian , K. 1984 . “ The origin of pterosaurs ” . In Proceedings of the Third Symposium on Mesozoic Terrestrial Ecosystems, Short Papers , Edited by: Reif , W.-E. and Westphal , F. 163 – 168 . Tübingen : Attempto Verlag .
- Padian , K. 1985 . The origin and aerodynamics of flight in extinct vertebrates . Palaeontology , 28 : 413 – 433 .
- Padian , K. 1987 . “ The case of the bat-winged pterosaur ” . In Dinosaurs Past and Present , Edited by: Czerkas , S.J. and Oison , E.C. Vol. II , 65 – 81 . California/ Seattle : Natural History Museum of Los Angeles County/University of Washington Press .
- Padian , K. 1991 . “ Pterosaurs: were they functional birds or functional bats? ” . In Biomechanics in Evolution , Edited by: Rayner , J.M.V. and Wooton , R.J. 145 – 160 . Cambridge : Cambridge University Press .
- Padian , K. and Rayner , J.V. 1993 . The wings of pterosaurs . American Journal of Science , 239-A : 91 – 166 .
- Peabody , FE. 1948 . Reptile and amphibian trackways from the Moenkopi Formation of Arizona and Utah . University of California Publications, Bulletin of the Department of Geological Sciences , 27 (8) : 325 – 340 .
- Pennycuick , C.J. 1986 . “ Mechanical constraints on the evolution of flight ” . In The Origin of Birds and the Evolution of Flight , Edited by: Padian , K. Vol. 8 , 83 – 98 . San Francisco : Memoirs of the California Academy of Sciences .
- Pennycuick , C.J. 1988 . On the reconstruction of pterosaurs and their manner of flight, with notes on vortex wakes . Biological Reviews , 63 : 299 – 331 .
- Peters , D. 2000a . Description and interpretation of interphalangeal lines in tetrapods . Ichnos , 7 : 11 – 41 .
- Peters , D. 2000b . A redescription of four prolacertiform genera and implications for pterosaur phylogenesis . Rivista Italiana di Paleontologia e Stratigrafia , 106 : 293 – 336 .
- Peters , S.D. and Gutmann , W.F. 1985 . “ Constructional and functional preconditions for the transition to powered flight in vertebrates ” . In The Beginning of Birds. Proceedings of the International Archaeopteryx Conference, Eichstatt 1984 , Edited by: Hecht , M. , Ostrom , J.H. , Viohl , G. and Wellnhofer , P. 239 – 242 . Eichstatt : Freundes Jura Museum .
- Reisz , R. and Sues , H-D. 2001 . Longisquama does not have feathers . Journal of Vertebrate Paleontology , 21 (Supplement to No. 3) : 92A
- Renesto , S. 1994 . A new prolacertiform reptile from the Late Triassic of Northern Italy . Rivista Italiana di Paleontologia e Stratigrafia , 100 : 285 – 306 .
- Romer , A.S. 1956 . Osteology of (he Reptiles , Chicago : University of Chicago Press .
- Romer , A.S. 1972 . The Chanares (Argentina) Triassic reptile fauna. XIII An early ornithosuchid pseudosuchian, Gradlisuchus stipanicicorum, gen. et sp. nov. . Breoiora , 389 : 1 – 24 .
- Russell , A.P. and Bauer , A.M. 1992 . The m. caudifemoralis longus and its relationship to caudal autotomy and locomotion in lizards (Reptilia: Sauria) . Journal of the Zoological Society of London , 227 : 127 – 143 .
- Schaller , D. 1985 . “ Wing Evolution ” . In The Beginning of Birds. Proceedings of the International Archaeopteryx Conference, Eichstätt 1984 , Edited by: Hecht , M. , Ostrom , J.H. , Viohl , G. and Wellnhofer , P. 333 – 348 . Eichstätt : Freundes Jura Museum .
- Serene , PC. 1991 . Basal archosaurs: phylogenetic relationships and functional implications. Society of Vertebrate Paleontology, Memoir 2 . Journal of Vertebrate Paleontology , 21 (Supplement to No. 4) : 1 – 53 .
- Sereno , P.C. 1996 . The phylogenetic position of pterosaurs within Archosauria . Journal of Vertebrate Paleontology , 16 (Supplement to No. 3) : 65A
- Sereno , P.C. and Wild , R. 1992 . Procompsognathus: theropod, "thecodont" or both? . Journal of Vertebrate Paleontology , 12 : 435 – 458 .
- Sereno , P.C. and Arcucci , A. 1994 . Dinosaurian precursors from the Middle Triassic of Argentina: Marasuchus lilloensis, gen. nov. . Journal of Vertebrate Paleontology , 14 : 53 – 73 .
- Sharov , A.G. 1970 . Unusual reptile from the Lower Triassic of Fergana . Paleontological Zhurnal 1970 , : 127 – 131 . (In Russian)
- Sharov , A.G. 1971 . New flying reptiles from the Mesozoic of Kazakhstan and Kirghizia . Akademia Nauk, Paleontological Institute, Moscow, Trudy , 130 : 104 – 113 . (In Russian)
- Shine , R. and Lambeck , R. 1989 . Ecology of frillneck lizards Chlamydosaurus kingii (Agamidae) in tropical Australia . Australian Wildlife Research , 16 : 491 – 500 .
- Snyder , R. 1954 . The anatomy and function of the pelvic girdle and hindlimb in lizard locomotion . American Journal of Anatomy , 95 : 1 – 36 .
- Stein , R.S. 1975 . Dynamic analysis of Pteranodon ingens: a reptilian adaptation of flight . Journal of Paleontology , 49 : 534 – 548 .
- Stokes , W.L. 1957 . Pterodactyl tracks from the Morrison Formation . Journal of Paleontology , 31 : 952 – 954 .
- Stromer , E. 1910 . Bemerkungen zur Rekonstruktion eines Flugsaurier-Skelettes . Monatsberichte der deutschen Geologischen Gesellschaft , 62 : 85 – 91 .
- Stromer , E. 1913 . Rekonstruktion des flugsauriers Rhamphorhynchus gemminigi H.v.M. . Neues Jahrbuch für Mineralogie, Geologie und Paläontologie , 2 : 49 – 68 .
- Unwin , D.M. 1987 . Pterosaur locomotion: joggers or waddlers? . Nature , 327 : 13 – 14 .
- Unwin , D.M . 1988 . New remains of the pterosaur Dimorphodon (Pterosauria: Rhamphorhynchoidea) and the terrestrial locomotion of early pterosaurs . Modem Geology , 13 : 57 – 68 .
- Unwin , D.M. and Bakhurina , N.N. 1994 . Sardes pilosus and the nature of the pterosaur flight apparatus . Nature , 371 : 62 – 64 .
- Unwin , D.M. , Frey , E. , Martill , D.M , Clarke , J.B. and Riess , J. 1996 . On the nature of the pteroid in pterosaurs . Proceedings of the Royal Society, London, B , 263 : 45 – 52 .
- Wang , X. , Zhou , Z. , Zhang , E and Xu , X. 2002 . A nearly completely articulated rhamphorhynchoid pterosaur with exceptionally well-preserved wing membranes and "hairs" from Inner Mongolia, northeast China . Chinese Science Bulletin , 47 (3) : 226 – 230 .
- Wellnhofer , P. 1975 . Die Rhamphorhynchoidea (Pterosauria) der Oberjura-Plattenkalke Süddeutschlands . Palaeontographica A (Stuttgart) , 148 : 1 – 33 . (148), 132-186 and (149), 1-30
- Wellnhofer , P. 1978 . Pterosauria. Handbuch der Paläoherpetologie , 1 – 82 . Stuttgart : Gustav Fischer Verlag . Teil 19
- Wellnhofer , P. 1987 . Die Flughaut von Pterodactylus (Reptilia: Pterosauria) am Beispiel des Wieners Exemplares von Pterodactylus kochi (Wagner) . Annalen des Naturhistorischen Museums Wien , 88A : 149 – 162 .
- Wellnhofer , P. 1991 . The Illustrated Encyclopedia of Pterosaurs , London : Salamander Books .
- Wdd , R. 1973 . Die Triasfauna der Tessiner Kalkalpen XXIII Tanystropheus longobardicus (Bassani) (Neue Ergebnisse) . Schweizerische Paläontologische Abhandlungen , 95 : 1 – 162 .
- Wild , R. 1978 . Die Flugsaurier (Reptilia, Pterosauria) aus der Oberen Trias von Cene bei Bergamo, Italien . Bolletino della Societa Paleontolpgica Italiana , 17 : 176 – 256 .
- Wild , R. 1983 . “ Über den Ursprung der Flugsaurier ” . In Y/eltenberger Akademie, Erwin Rutte-Festschrift , 231 – 238 . Weltenburg : Kelheim .
- Wild , R. 1984 . A new pterosaur (Reptilia, Pterosauria) from the Upper Triassic (Norian) of Friuli, Italy . Gortania, Atti Museo Friulano di Storia Naturale , 5 : 45 – 62 .
- Wild , R. 1993 . A juvenile specimen of Eudimorphodon ranzii Zambelli (Reptilia, Pterosauria) from the Upper Triassic (Norian) of Bergamo. Rivista Museo Civco di Scienze Naturali "E . Caffi" Bergamo , 16 : 95 – 120 .
A New Model for the Evolution of the Pterosaur Wing--with a twist
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