Hyaenidae (Carnivora) from the Late Miocene hominid locality of Hammerschmiede (Bavaria, Germany)

References

• Alba DM, Moyà-Solà S, Casanovas-Vilar I, Galindo J, Robles JM, Rotgers C, Furió M, Angelone C, Köhler M, Garcés M, et al. 2006. Los vertebrados fósiles del Abocador de Can Mata (els Hostalets de Pierola, l’Anoia, Cataluña), una sucesión de localidades del Aragoniense superior (MN6 y MN7+8) de la cuenca del Vallès-Penedès. Campañas 2002-2003, 2004 y 2005. Estudios Geológicos. 62(1):295–312. Spanish. doi:10.3989/egeol.0662127.
   Google Scholar
• Andrews CW. 1918. Note on some fossil mammals from Salonica and Imbros. Geol Mag. 6:540–543. doi:10.1017/S0016756800201076.
   Google Scholar
• Andrews P, Whybrow P, Stringer C. 1980. Stratigraphy and Palaeontology of Miocene deposits at Yeni Eskihisar, Turkey. Newsletters on Stratigraphy. 9(1):49–57. doi:10.1127/nos/9/1980/49.
   Google Scholar
• Antón M, Turner A, Salesa M, Morales J. 2006. A complete skull of Chasmaporthetes lunensis (Carnivora, Hyaenidae) from the Spanish Pliocene site of La Puebla de Valverde (Teruel). Estudios Geológicos. 62(1):375–388. doi:10.3989/egeol.0662132.
   Google Scholar
• Bastl K, Nagel D, Morlo M, Göhlich UB. 2020. The Carnivora (Mammalia) from the middle Miocene locality of Gračanica (Bugojno Basin, Gornji Vakuf, Bosnia And Herzegovina). Palaeobiodiversity and Palaeoenvironments. 100(2):307–319. doi:10.1007/s12549-018-0353-0.
   Web of Science ®Google Scholar
• Beke D 2010. The fossil cave hyena of Goyet, Walsin and Hastière (Belgium): osteometry and taphonomy [master’s thesis, MSc Thesis]. Gent: Universiteit Gent.
   Google Scholar
• Böhme M, Spassov N, DeSilva J, Begun DR. 2020. Reply to: reevaluating bipedalism in Danuvius. Nature. 586(7827):E4–E5. doi:10.1038/s41586-020-2737-3.
   PubMed Web of Science ®Google Scholar
• Böhme M, Spassov N, Fuss J, Tröscher A, Deane A, Prieto A, Kirscher U, Lechner T, Begun D. 2019. A new Miocene ape and locomotion in the ancestor of great apes and humans. Nature. 575:489–493. doi:10.1038/s41586-019-1731-0.
   PubMed Web of Science ®Google Scholar
• Böhme M, Spassov N, Majidifard MR, Gärtner A, Kirscher U, Marks M, Dietzel C, Uhlig G, El Afty H, Begun DR, et al. 2021. Neogene hyperaridity in Arabia frove unidirectional mammalian dispersal between Africa and Eurasia. Nature Communications Earth & Environment. 2:1–13.
   Google Scholar
• Brisson AD. 1762. Le regne animal divisé en IX classes. Paris: Jean-Baptiste Bauche. French.
   Google Scholar
• Coca-Ortega C, Pérez-Claros JA. 2019. Characterizing ecomorphological patterns in hyenids: a multivariate approach using postcanine dentition. PeerJ. 6:e6238. doi:10.7717/peerj.6238.
   PubMed Web of Science ®Google Scholar
• Colbert EH. 1939. Carnivora of the Tung Gur Formation of Mongolia. Bulletin of the American Museum of Natural History. 76:47–8l.
   Google Scholar
• Crusafont Pairó M, Golpe Posse J. 1981. Hallazgo de una nueva especie del género Semigenetta (S. grandis) del Vindoboniense terminal de Castell de Barberà (Depresión prelitoral catalana). Boletin de la Real Sociedad Española de Historia Natural, Sección Geológica. 79:67–76.
   Google Scholar
• Crusafont Pairó M, Golpe Posse M. 1973. Nuevos hallazgos de Progenetta montadai en el Miocene de Cataluna. Boletin Geológico y Minero. 84: 105–113. Spanish.
   Google Scholar
• Crusafont Pairó M, Kurtén B. 1976. Bears and Bear-Dogs from the Vallesian of the Vallés-Penedés Basin, Spain. Acta Zoologica Fennica. 144:1–29.
   Google Scholar
• Crusafont Pairó M, Petter C. 1969. Contribution a I’ etude des Hyaenidae. La sous-familie des Ictitheriinae. Annales de Paléontologie (Vertébrés). 55: 89–121. French.
   Google Scholar
• de Beaumont C. 1964. Note sur la région otique d’ Ictitherium hipparionum (Gervais) (Carnivora). Archives des Sciences, Genève. 17:339–342.
   Google Scholar
• de Beaumont C. 1986. Les Carnivores (Mammiferes) du Néogène de Höwenegg/ Hegau,Baden-Wurttemberg. Carolinea. 44:35–45.
   Google Scholar
• de Bonis L. 2005. Carnivora (Mammalia) from the late Miocene of Akkaşdağı, Turkey. Geodiversitas. 27(4):567–589.
   Web of Science ®Google Scholar
• Depéret C. 1892. La faune des mammifères miocènes de La Crive St-Alban. Archives Du Museum, Lyon. 5: 1–89. French.
   Google Scholar
• Erxleben JCP. 1777. Systema regni animalis per classes, ordines, genera, species, varietates cvm synonymia et historia animalivm. Leipzig: Weigand, Leipzig.
   Google Scholar
• Filhol H. 1890. Étude sur les mammifères de Sansan. Bibliothèque des Hautes Études section des sciences naturelles. 37(1):1–319. French.
   Google Scholar
• Forsyth Major CI. 1903. New Carnivora from the middle Miocene of La Grive St-Alban. Geological Magazine N S. 1:534–538. doi:10.1017/S0016756800115365.
   Google Scholar
• Gabunia L. 1973. Fossil vertebrates of the fauna of Bjelometschskaya. Tbilisi: Akademii Nauk, Georgian Soviet Socialist Republic. Russian.
   Google Scholar
• Gaudry A. 1861. Note sur les carnassiers fossiles de Pikermi (Grèce). Bulletin de la Société Géologique de France. 18:527–538.
   Google Scholar
• Geoffroy Saint-Hilaire I. 1824. Description d’un nouveau genre de mammifères carnassiers sous le nom de Protèle. Memoires du Museum d ‘Histoire Naturelle, Paris. 11: 354–371. French.
   Google Scholar
• Gervais P. 1846. Observations sur diverses especes de mamiferes fossiles du midi de France. Annales des Sciences Naturelles, ser 3. 5: 248–265. French.
   Google Scholar
• Gervais P. 1850. Zoologie et Paleontologie Françaises. Paris: Atthus Bertrand.
   Google Scholar
• Ginsburg L. 1999. Order Carnivora. In: Rössner GE, Heissig K, editors. The Miocene Land Mammals of Europe. Munich: Friedrich Pfeil; p. 109–148.
   Google Scholar
• Gray JE. 1821. On the natural arrangement of vertebrose animals. London Medical Repository. 15:296–310.
   Google Scholar
• Hay OP. 1921. Descriptions of species of Pleistocene Vertebrata, types or specimens most of which are preserved in the United States National Museum. Proceedings of the United States National Museum. 59:599–642. doi:10.5479/si.00963801.59-2391.599.
   Google Scholar
• Hendey QB. 1974. New fossil carnivores from the Swartkrans australopithecine site (Mammalia: carnivora). Annals of the Transvaal Museum. 29:27–48.
   Google Scholar
• Hensel RF. 1862. Über die Reste einiger Säugetierarten von Pikermi in der Münchener Sammlung. Monatsberichte der Akademie der Wissenschaften. 27:560–569.
   Google Scholar
• Howell FC. 1987. Preliminary Observations on Carnivora from the Sahabi Formation (Libya). In: Boaz N, El-Arnauti A, Gaziry AW, de Heinzelin J, Dechant Boaz D, editors. Neogene paleontology and geology of Sahabi. New York: A.R. Liss; p. 153–181.
   Google Scholar
• Howell FC, Petter G. 1985. Comparative observations on some Middle and Upper Miocene hyaenid genera: percrocuta Kretzoi, Allohyaena Kretzoi, Adcrocuta Kretzoi (Mammalia, Carnivora, Hyaenidae). Geobios. 18:419–476. doi:10.1016/S0016-6995(85)80001-2.
   Web of Science ®Google Scholar
• Kargopoulos N, Kampouridis P, Lechner T, Böhme M. 2021a. A review of Semigenetta (Viverridae, Carnivora) from the Miocene of Eurasia based on material from the hominid locality of Hammerschmiede (Germany). Geobios. doi:10.1016/j.geobios.2021.07.001
   Web of Science ®Google Scholar
• Kargopoulos N, Valenciano A, Kampouridis P, Lechner T, Böhme M. 2021b. An otter’s journey: a new species of Vishnuonyx (Carnivora, Lutrinae) from the hominid locality of Hammerschmiede (early Late Miocene; Bavaria, Germany) and the first report of the genus in Europe. Journal of Vertebrate Paleontology. doi:10.1080/02724634.2021.1948858
   Web of Science ®Google Scholar
• Kaup JJ. 1828. Hyaena. Isis Von Oken. 21(11):1144–1145.
   Google Scholar
• Kaya T, Geraads D, Tuna V. 2003. A new Middle Miocene mammalian fauna from Mordoğan (Western Turkey). Paläontologische Zeitschrift. 77(2):293–302. doi:10.1007/BF03006943.
   Google Scholar
• Kirscher U, Prieto J, Bachtadse V, Abdul Aziz H, Doppler G, Hagmaier M, Böhme M. 2016. A biochronologic tie-point for the base of the Tortonian stage in European terrestrial settings: magnetostratigraphy of the topmost Upper Freshwater Molasse sediments of the North Alpine Foreland Basin in Bavaria (Germany). Newsletters on Stratigraphy. 49(3):445–467. doi:10.1127/nos/2016/0288.
   Web of Science ®Google Scholar
• Koufos G. 1979. Preliminary study on the mammalian fauna (Carnivora - Proboscidea, Perissodactyla) of Axios Valley (Macedonia - Greece). Annales Géologiques des Pays Helléniques, Hors Serie. 1979:631–635.
   Google Scholar
• Koufos G. 1995. The late Miocene percrocutas (Carnivora, Mammalia) of Macedonia, Greece. Palaeovertebrata. 24:67–84.
   Google Scholar
• Koufos G. 2000. Revision of the late Miocene carnivores from the Axios valley, Macedonia, Greece. Münchner Geowissenschaftliche Abhandlungen A. 39:51–92.
   Google Scholar
• Koufos G. 2011. The Miocene carnivore assemblage of Greece. Estudios Geológicos. 67(2):291–320. doi:10.3989/egeol.40560.190.
   Web of Science ®Google Scholar
• Koufos G. 2012. New material of Carnivora (Mammalia) from the Late Miocene of Axios Valley, Macedonia, Greece. Comptes Rendus Palevol. 11(1):49–64. doi:10.1016/j.crpv.2011.09.004.
   Web of Science ®Google Scholar
• Koufos G, Mayda S, Kaya T. 2018. New carnivoran remains from the Late Miocene of Turkey. PalZ. 92(1):131–162. doi:10.1007/s12542-017-0376-2.
   Web of Science ®Google Scholar
• Kretzoi M. 1938. Die Raubtiere von Gombaszög nebst einer Obersicht der Gesamtfauna. Annales Museum Nationale Hungaricum. 31: 89–157. German.
   Google Scholar
• Kurtén B. 1982. Status of the Fossil Hyaenids Ictitherium viverrinum and Thalassictis robusta (Mammalia). Z Geol Wiss Berlin. 10(7):1009–1018.
   Web of Science ®Google Scholar
• Lartet E. 1851. Notice sur la colline de Sansan, suivie d’une recapitulation des diverses espèces d’animaux vertébrés fossiles, trouvés soit à Sansan, soit dans d’autres gisements du terrain tertiaire miocène dans le bassin souspyrénéen. Auch: J. A. Portes. French.
   Google Scholar
• Linnaeus C. 1758. Systema naturae per regna tria naturae, secundum classis, ordines, genera, species cum characteribus, differentiis, synonymis, locis. 10th ed. Stockholm: Laurentius Salvius. Latin.
   Google Scholar
• Lungu A 1978. The Hipparion fauna of the middle Sarmatian of Moldova (Carnivores). Shtiintsa, Kishinev, 133 p. (in Russian).
   Google Scholar
• Lungu A, Rzebik-Kowalska B. 2011. Faunal assemblages, stratigraphy and taphonomy of the Late Miocene localities in the Republic Of Moldova. Krakow: Institute of Systematics and Evolution of Animals, Polish Academy of Sciences.
   Google Scholar
• Mayda S, Koufos G, Kaya T, Gul A. 2015. New carnivore material from the Middle Miocene of Turkey. Implications on biochronology and palaeoecology. Geobios. 48(1):9–23. doi:10.1016/j.geobios.2014.11.001.
   Web of Science ®Google Scholar
• Mayr H, Fahlbusch V. 1975. Eine unterpliozäne Kleinsäuger-fauna aus der Oberen Süßwasser-Molasse Bayerns. Mitteilungen der Bayerischen Staatssammlung für Paläontologie und historische Geologie. 15:91–111.
   Google Scholar
• Morlo M. 1997. Die Raubtiere (Mammalia, Carnivora) aus dem Turolium von Dorn-Dürkheim 1 (Rheinhessen). Teil 1: mustelida, Hyaenidae, Percrocutidae, Felidae. Courier Forschungs-Institut Seckenberg. 197:11–47.
   Google Scholar
• NOW Community. 2021. New and Old Worlds Database of Fossil Mammals (NOW). Licensed under CC BY 4.0. [accessed 2021 Oct 18]. http://www.helsinki.fi/science/now/.
   Google Scholar
• Ozansoy F. 1961. Hyaena senyureki Ozansoy du Pliocène supérieur (Sinap moyen) et la repartition vertical du genre Hyaena dans le système pliocène de Turqie. Bulletin of Mineral Research and Exploration. 57: 97–120. French.
   Google Scholar
• Ozansoy F. 1965. Etudes des gisements continentaux et des mammifères du Cénozoique de Turquie. Mémoires de la Société géologique de France, N S 44. 102: 1–92. French.
   Google Scholar
• Pavlović M, Thenius E. 1965. Eine neue Hyäne (Carnivora, Mammalia) aus dem Miozän Jugoslawiens und ihre phlyogenetische Stellung. Anzeiger der Österreichischen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse. 102(2):177–185.
   Google Scholar
• Pavlow M. 1908. Quelques carnivores fossiles du gouvernement de Kherson et de Bessarabie. Memoires de la Societe des Naturalistes de la Nouvelle-Russie, Odessa. 32:27–48.
   Google Scholar
• Peigné S. 2016. Carnivora. Geodiversitas. 38(2):197–224. doi:10.5252/g2016n2a4.
   Web of Science ®Google Scholar
• Pilgrim GE. 1910. Notices of new mammalian genera and species from the Tertiaries of India. Records of the Geological Survey of India. 40:63–71.
   Google Scholar
• Pilgrim GE. 1931. Catalogue of the Pontian Carnivora of Europe in the Department of Geology. London: British Museum (Natural History).
   Google Scholar
• Pilgrim GE. 1932. The fossil Carnivora of India. Paleontologica Indica, n s. 18:1–232.
   Google Scholar
• Radović P, Mayda S, Alaburić S, Marković Z. 2021. Percrocuta miocenica (Percrocutidae, Carnivora) from the middle Miocene of Brajkovac (Central Serbia). Geobios. 65:41–49. doi:10.1016/j.geobios.2021.02.001.
   Web of Science ®Google Scholar
• Robles JM 2014. Miocene Carnivorans from the Valles-Penedes Basin (NE Iberian Peninsula) [dissertation]. Barcelona: Universitat Autonoma de Barcelona.
   Google Scholar
• Roth J, Wagner A. 1854. Die fossilen Knocheniiberreste von Pikermi in Griechenland. Abhandlungen der Bayerisehen Akademie der Wissensehaften. 7(37):1–464.
   Google Scholar
• Schlosser M. 1903. Die Fossilen Säugethiere Chinas nebst einer Odontographie der recenten Antilopen. Abhandlungen der Bayerischen Akademie derWissenschaften. 22:1–221.
   Google Scholar
• Schmidt-Kittler N. 1976. Raubtiere aus dem Jungtertiar Kleinasiens. Palaeontographica Abt A. 155:1–131.
   Google Scholar
• Semenov Y. 1988. Ictitherium spelaeum novyj vid iktiteriya (Carnivora, Viverridae) iz srednego sarmata Ukrainy. [Ictitherium spelaeum sp. n. (Carnivora, Viverridae) from Middle Sarmat of the Ukraine]. Vestnik Zoologii. 1988: 45–49. (In Russian, English abstract).
   Google Scholar
• Semenov Y 1989. Iktiterii i morfologieheski skhodnye hieny neogena SSSR [Ictitheres and morphologically related hyaenas from the Neogene of the USSR] [dissertation]. Kiev: Naukova Dumka. (In Russian, English summary).
   Google Scholar
• Semenov Y. 1994. Allohyaena sarmatica (Carnivora, Mammalia) – a new hyaenid species from the late Miocene of the Ukraine. Acta Zoologica Cracoviensia. 37(1):31–38.
   Google Scholar
• Semenov Y. 2008. Taxonomical reappraisal of “ictitheres” (Mammalia, Carnivora) from the Late Miocene of Kenya. C R Palevol. 7(8):529–539. doi:10.1016/j.crpv.2008.09.012.
   Web of Science ®Google Scholar
• Solounias N, de Beaumont G. 1981. Brief remarks on certain large-sized Turolian hyaenids (Mammals, Carnivores) from Pikermi and Samos, Greece. Archives des Sciences, Genève. 34:293–304.
   Google Scholar
• Sparrman A. 1783. Resa till Goda Hopps-Udden Sodra Polkretsen och Omk: ringJordklotet samt till Hottentott-och Caffer-landen aren 1722-76. Stockholm: AJ Mordstrom.
   Google Scholar
• Spassov N, Koufos G. 2002. The first appearance of Dinocrocuta gigantea and Machairodus aphanistus (Mammalia, Carnivora) in the Miocene of Bulgaria. Mitt Bayer Staatsslg Paläont Hist Geol. 42:83–101.
   Google Scholar
• Thenius E. 1966. Zur Stammesgeschichte der Hyänen (Carnivora, Mammalia). Zeitschrift fur Säugetierkunde. 31:293–300.
   Google Scholar
• Thunberg CP. 1820. Beskrifning och Techning pa ett nytt species Hyaena brunnea. Stockholm: kungl. Svenska Vetenskapsakademiens Handlingar Ser 3. 8:59–65.
   Google Scholar
• Torre D. 1989. Plioviverrops faventinus n. sp., a new carnivore of late Messinian age. Bolletino della Società Paleontologica Italiana. 28(2–3):323–327.
   Google Scholar
• Trouessart EL. 1897. Catalogus Mammalium tam Vivientum quam Fossilium. Berlin: Berolini,R. Friedländer & Sohn. Latin.
   Google Scholar
• Tseng ZJ, Li Q, Wang X. 2013. A new cursorial hyena from Tibet, and analysis of biostratigraphy, paleozoogeography, and dental morphology of Chasmaporthetes (Mammalia, Carnivora). Journal of Vertebrate Paleontology. 33(6):1457–1471. doi:10.1080/02724634.2013.775142.
   Web of Science ®Google Scholar
• Turner A, Antón M, Werdelin L. 2008. Taxonomy and evolutionary patterns in the fossil Hyaenidae of Europe. Geobios. 41(5):677–687. doi:10.1016/j.geobios.2008.01.001.
   Web of Science ®Google Scholar
• van Dam JA. 1997. The small mammals from the upper Miocene of the Teruel-Alfambra region (Spain): paleobiology and paleoclimatic reconstructions. Geologica Ultraiectina. 156:1–204.
   Google Scholar
• Vangengeim E, Tesakov A. 2013. Late Miocene Mammal Localities of Eastern Europe and Western Asia: toward Biostratigraphic Synthesis. In: Wang X, Flynn L, Fortelius M, editors. Fossil Mammals of Asia. New York: Columbia University Press; p. 521–537.
   Google Scholar
• Villalta Comella JF, Crusafont Pairó M. 1943. Los Vertebrados del Mioceno Continental de la Cuenca Valles-Panades (Provincia de Barcelona). I Insectivoros. - II. Carnivoros. Boletin del Instituto Geológico y Minero de España. 56:147–192.
   Google Scholar
• Vinuesa V, Madurell-Malapeira J, Werdelin L, Robles JM, Obradó P, Alba D. 2017. A new skull of Hyaenictis Gaudry, 1861 (Carnivora: hyaenidae) shows incipient adaptations to durophagy. J Mamm Evol. 24(2):207–219. doi:10.1007/s10914-016-9334-0.
   Web of Science ®Google Scholar
• Viranta S, Werdelin L. 2003. Carnivora. In: Fortelius M, Kappelman J, Sen S, Bernor R, editors. Geology and Paleontology of the Miocene Sinap Formation, Turkey. New York: Columbia University Press; p. 178–193.
   Google Scholar
• Viret J. 1951. Catalogue critique de la faune des mammiferes Miocenes de La Grive Saint-Alban (Isere). Nouvelles Archives du Museum d ‘Histoire Naturelle, Lyon. 3:1–104.
   Google Scholar
• Wagner JA. 1848. Urweltliche Säugethiere-Überreste aus Griechenland. Abhandlungen der Bayerischen Akademie der Wissenschaften. 5:335–378.
   Google Scholar
• Wang X, Tedford R. 2008. Dogs: their fossil relatives and evolutionary history. New York (NY): Columbia University Press.
   Google Scholar
• Wang X, Ye J, Meng J, Wu W, Liu LP, Bi S. 1998. Carnivora from Middle Miocene of Northern Junggar Basin, Xinjiang Autonomous Region, China. Vertebrata PalAsiatica. 36(3):218–243.
   Google Scholar
• Werdelin L. 1988. Studies of fossil hyaenas: the genera Thalassictis Gervais ex Nordmann, Palhyaena Gervais, Hyaenictitherium Kretzoi, Lycyaena Hensel and Palinhyaena Qiu, Huang & Guo. Zool J Linn Soc. 92(3):211–265. doi:10.1111/j.1096-3642.1988.tb01512.x.
   Web of Science ®Google Scholar
• Werdelin L. 1991. Hyaenas: the rise and fall of a carnivore family. Geologiska Föreningen I Stockholm Förhandlingar. 113(1):95. doi:10.1080/11035899109453840.
   Google Scholar
• Werdelin L. 2005. The Carnivora of Rudabánya (MN 9, Late Miocene, Hungary). Palaeontographica Italica. 90:163–180.
   Google Scholar
• Werdelin L, Solounias N. 1990. Studies of fossil hyaenids: the genus Adcrocuta Kretzoi and the interrelationships of some hyaenid taxa. Zool J Linn Soc. 98(4):363–386. doi:10.1111/j.1096-3642.1990.tb01206.x.
   Web of Science ®Google Scholar
• Werdelin L, Solounias N. 1991. The Hyaenidae: taxonomy, systematics and evolution. Fossils and Strata. 30:1–104.
   Google Scholar
• Xiong W-Y. 2019. Basicranial morphology of Late Miocene Dinocrocuta gigantea (Carnivora: hyaenidae) from Fugu, Shaanxi. Vertebrata PalAsiatica. 57(4):274–307.
   Google Scholar
• Zdansky O. 1924. Jungtertiäre Carnivoren Chinas. Palaeontologia Sinica C. 2(1):1–223.
   Google Scholar
• Zhang Z, Xiangxu X 1996. New materials of Dinocrocuta gigantea found in Fugu
   Google Scholar