Concurrent evidence from ichnology and anatomy: the scelidotheriine ground sloths (Xenarthra, Folivora) from the Pleistocene of Argentina

References

• Alexander RM. 1989. Optimization and gaits in the locomotion of vertebrates. Physiol Rev. 69(4):1199–1227. doi:10.1152/physrev.1989.69.4.1199.
   PubMed Web of Science ®Google Scholar
• Amson E, Argot C, McDonald HG, De Muizon C. 2015. Osteology and functional morphology of the hind limb of the marine sloth Thalassocnus (Mammalia, Tardigrada). J Mamm Evol. 22(3):355–419. doi:10.1007/s10914-014-9274-5.
   Web of Science ®Google Scholar
• Aramayo SA. 2007. Neogene vertebrate palaeoichnology of the north Atlantic coast of the Río Negro province, Argentina. Arquivos do Museu Nacional. 65(4):573–584.
   Google Scholar
• Aramayo SA, and Manera de Bianco T. 1987. Hallazgo de una icnofauna continental (Pleistoceno tardío) en la localidad de Pehuén-Có (Partido de Coronel Rosales), Provincia de Buenos Aires, Argentina. Parte II. Carnivora, Artiodactyla y Aves. P4° Congreso Latinoamericano de Paleontología, La Paz, Bolivia. Resúmenes; p. 532–547.
   Google Scholar
• Aramayo SA, Manera de Bianco T. 1996. Edad y nuevos hallazgos de icnitas de mamíferos y aves en el yacimiento paleoicnologico de Pehuen-Co (Pleistoceno tardio), provincia de Buenos Aires, Argentina. Publicación Electrónica de la Asociación Paleontológica Argentina. 4(1):47–58.
   Google Scholar
• Aramayo SA, Manera de Bianco T, Bastianelli NV, Melchor RN. 2015. Pehuen Co: updated taxonomic review of a late Pleistocene ichnological site in Argentina. Palaeogeogr Palaeocl. 439:144–165. doi:10.1016/j.palaeo.2015.07.006.
   Web of Science ®Google Scholar
• Bargo MS, Vizcaíno SF, Archuby FM, Blanco RE. 2000. Limb bone proportions, strength and digging in some Lujanian (Late Pleistocene-Early Holocene) mylodontid ground sloths (Mammalia, Xenarthra). J Vertebr Paleontol. 20(3):601–610. doi:10.1671/0272-4634(2000)020[0601:LBPSAD]2.0.CO;2.
   Web of Science ®Google Scholar
• Bennett MR, Bustos D, Belvedere M, Martinez P, Reynolds SC, Urban T. 2019. Soft-sediment deformation below mammoth tracks at white sands national monument (New Mexico) with implications for biomechanical inferences from tracks. Palaeogeogr Palaeocl. 527:25–38. doi:10.1016/j.palaeo.2019.04.023.
   Web of Science ®Google Scholar
• Blanco RE, Czerwonogora A. 2003. The gait of Megatherium Cuvier 1796. Senckenberg Biol. 83:61–68.
   Google Scholar
• Bonaparte JF. 1965. Nuevas icnitas de la Quebrada del Yeso (La Rioja) y reconsideración de la edad de los afloramientos. Acta Geológica Lilloana. 7:5–16.
   Google Scholar
• Boscaini A, Pujos F, Gaudin TJ. 2019. A reappraisal of the phylogeny of Mylodontidae (Mammalia, Xenarthra) and the divergence of mylodontine and lestodontine sloths. Zool Scr. 48(6):691–710. doi:10.1111/zsc.12376.
   Web of Science ®Google Scholar
• Bustos D, Jakeway J, Urban TM, Holliday VT, Fenerty B, Raichlen DA, Marcin B, Reynolds SC, Allen BD, Love DW, et al. 2018. Footprints preserve terminal Pleistocene hunt? Human-sloth interactions in North America. Science Advances. 4(4):eaar7621. doi:10.1126/sciadv.aar7621.
   PubMed Web of Science ®Google Scholar
• Camens A, Wells R. 2009. Diprotodontid footprints from the Pliocene of Central Australia. J Vertebr Paleontol. 29(3):863–869. doi:10.1671/039.029.0316.
   Web of Science ®Google Scholar
• Carey SP, Camens AB, Cupper ML, Grün R, Hellstrom JC, McKnight SW, McIennan I, Pickering DA, Trusler P, Aubert M. 2011. A diverse Pleistocene marsupial trackway assemblage from the Victorian Volcanic Plains, Australia. Quaternary Sci Rev. 30(5–6):591–610. doi:10.1016/j.quascirev.2010.11.021.
   Web of Science ®Google Scholar
• Carrano MT, Wilson JA. 2001. Taxon distributions and the tetrapod track record. Paleobiology. 27(3):564–582. doi:10.1666/0094-8373(2001)027<0564:TDATTT>2.0.CO;2.
   Web of Science ®Google Scholar
• Casamiquela RM. 1974. El bipedismo de los megaterioideos estudio de pisadas fosiles en la Formación Rio Negro tipica. Ameghiniana. 11(3):249–282.
   Google Scholar
• Casamiquela RM. 1983. Pisadas del Pleistoceno (Superior?) del balneario de Monte Hermoso, Buenos Aires: la confirmación del andar bipedal en los megaterioideos. Cuadernos del Instituto Superior “Juan XXIII”. 4:2–21.
   Google Scholar
• Casinos A. 1996. Bipedalism and quadrupedalism in Megatherium: an attempt at biomechanical reconstruction. Lethaia. 29(1):87–96. doi:10.1111/j.1502-3931.1996.tb01842.x.
   Web of Science ®Google Scholar
• Coombs MC. 1983. Large mammalian clawed herbivores: a comparative study. T Am Philos Soc. 73(7):1–96. doi:10.2307/3137420.
   Web of Science ®Google Scholar
• Cuenca Anaya J 1992. El aparato locomotor de los escelidoterios (Edentata, Mammalia) y su paleobiología [PhD Dissertation]. Madrid: Universidad Complutense de Madrid.
   Google Scholar
• De Iuliis G 1996. A systematic review of the Megatheriinae (Mammalia: xenarthra: megatheriidae) [Ph.D. dissertation]. Canada: University of Toronto.
   Google Scholar
• de Toledo PM. 1996. Locomotory patterns within the Pleistocene sloths. PhD Dissertation. University of Colorado, USA, 316 pp.
   Google Scholar
• Diedrich CG. 2011. An overview of the ichnological and ethological studies in the Cave Bear Den in Urşilor Cave (Western Carpathians, Romania). Ichnos. 18(1):9–26. doi:10.1080/10420940.2011.552578.
   Web of Science ®Google Scholar
• Dillon A, Rabassa J. 1985. Miembro La Chumbiada, Formación Luján (Pleistoceno, provincia de Buenos Aires): una nueva unidad estratigráfica del valle del Río Salado. I Jornadas Geológicas Bonaerenses Actas. 1:1–27.
   Google Scholar
• Fariña RA, Vizcaíno SF, Bargo MS. 1998. Body mass estimations in Lujanian (late Pleistocene-early Holocene of South America) mammal megafauna. Mastozoología Neotropical. 5(2):87–108.
   Google Scholar
• Fariña RA, Vizcaíno SF, De Iuliis G, Tambusso S. 2013. Megafauna: giant Beasts of Pleistocene South America. Bloomington: Indiana University Press.
   Google Scholar
• Fidalgo F, De Francesco F, and Pascual R. 1975. Geología superficial de la llanura bonaerense (Argentina). 6° Congreso Geológico Argentino. Bahia Blanca, Argentina. Relatorio; p. 103–138.
   Google Scholar
• Fisher DC. 2018. Paleobiology of pleistocene proboscideans. Annu Rev Earth Pl Sc. 46(1):229–260. doi:10.1146/annurev-earth-060115-012437.
   Google Scholar
• Frenguelli J. 1950. Ichnites en el Paleozoico Superior del oeste argentino. Revista de la Asociación Geológica Argentina. 5(3):136–149.
   Google Scholar
• Gaudin TJ. 2004. Phylogenetic relationships among sloths (Mammalia, Xenarthra, Tardigrada): the craniodental evidence. Zool J Linn Soc-Lond. 140(2):255–305. doi:10.1111/j.1096-3642.2003.00100.x.
   Web of Science ®Google Scholar
• Harrold F, Abboud RJ. 2018. Biomechanics of the Foot and Ankle. In: Robinson A, Brodsky JW, Negrine JP, editors. Core Topics on Foot and Ankle Surgery. Cambridge: Cambridge University Press; p. 22–43.
   Google Scholar
• Hirschfeld SE. 1985. Ground sloths from the Friasian La Venta fauna, with additions to the Pre-Friasian Coyaima fauna of Colombia, South America. Berkeley: University of California Press.
   Google Scholar
• Hornung JJ, Böhme A, Schlüter N, Reich M. 2016. Diversity, ontogeny, or both? A morphometric approach to iguanodontian ornithopod (Dinosauria: ornithischia) track assemblages from the Berriasian (Lower Cretaceous) of North Western Germany. In: Marty D, Falkingham PL, Richter A, editors. Dinosaur Tracks: the Next Steps. Bloomington: Indiana University Press; p. 202–225.
   Google Scholar
• Krapovickas V, Ciccioli PL, Mángano MG, Marsicano CA, Limarino CO. 2009. Paleobiology and paleoecology of an arid–semiarid Miocene South American ichnofauna in anastomosed fluvial deposits. Palaeogeogr Palaeocl. 284(3–4):129–152. doi:10.1016/j.palaeo.2009.09.015.
   Web of Science ®Google Scholar
• Leonardi G. 1987. Glossary and manual of tetrapod footprint palaeoichnology. Brasilia: Ministério das Minas e Energia, Departamento Nacional da Produçao Mineral.
   Google Scholar
• Leonardi G. 1994. Annotated Atlas of South America tetrapod footprints (Devonian to Holocene) with an Appendix on Mexico and Central Americas. Rio de Janeiro: Ministério de Minas e Energía, Secretaría de Minas e Metalurgía.
   Google Scholar
• Lockley MG. 2011. The ichnotaxonomic status of Brasilichnium with special reference to occurrences in the Navajo Sandstone (Lower Jurassic) in the western USA. New Mexico Museum of Natural History and Science Bulletin. 53:306–315.
   Google Scholar
• Lockley MG, Lires J, García-Ramos JC, Pinuela L, Avanzini M. 2007. Shrinking the world’s largest dinosaur tracks: observations on the ichnotaxonomy of Gigantosauropus asturiensis and Hispanosauropus hauboldi from the Upper Jurassic of Asturias, Spain. Ichnos. 14(3–4):247–255. doi:10.1080/10420940601050048.
   Google Scholar
• Machado Cruvinel AC, Camargos de Britto Rosa M, Lopes Guimarães BL, Ferreira Amaral Silva C, de Sampaio da Silva PH, de Lima Borges Leão C, Peixoto JV, de Resende JJC, Cunha Lacreta JAC, Passamani M, et al. 2019. The lesser anteater (Tamandua tetradactyla) is not tetradactyla, but pentadactyla. Anat Histol Embryol. 48(3):256–263. doi:10.1111/ahe.12431.
   PubMed Web of Science ®Google Scholar
• Manera T, Bastianelli N. 2010. Nuevo registro de icnitas de mamíferos pleistocenos en Playa del Barco, Pehuen Co, provincia de Buenos Aires, Argentina. Paper presented at: Resúmenes del X Congreso Argentino de Paleontología y Bioestratigrafía-VII Congreso Latinoamericano de Paleontología. La Plata, Argentina.
   Google Scholar
• Marsh OC. 1883. On the Supposed Human Foot-prints recently found in Nevada. Am J Sci. 26(152):139. doi:10.2475/ajs.s3-26.152.139.
   Google Scholar
• Marty D, Falkingham PL, Richter A. 2016. Dinosaur track terminology: a glossary of terms. In: Marty D, Falkingham PL, Richter A, editors. Dinosaur Tracks: the Next Steps. Bloomington: Indiana University Press; p. 399–402.
   Google Scholar
• McDonald HG 1987. A systematic review of the Plio-Pleistocene scelidothere ground sloths (Mammalia: xenarthra, Mylodontidae) [PhD. Dissertation]. Canada, University of Toronto.
   Google Scholar
• McDonald HG. 2005. Paleoecology of extinct xenarthrans and the Great American Biotic Interchange. Bulletin Florida Museum Natural History. 45(4):313–333.
   Google Scholar
• McDonald HG. 2007. Biomechanical Inferences of Locomotion in Ground Sloths: integrating Morphological and Track Data. In: Lucas SG, Spielmann JA, Lockley MG, editors. Cenozoic Vertebrate Tracks and Traces. New Mexico: New Mexico Museum of Natural History and Science Bulletin; p. 201–208.
   Google Scholar
• McDonald HG. 2012. Evolution of the pedolateral foot in ground sloths: patterns of change in the astragalus. J Mamm Evol. 19(3):209–215. doi:10.1007/s10914-011-9182-x.
   Web of Science ®Google Scholar
• McDonald HG, De Iuliis G. 2008. Fossil history of sloths. In: Vizcaíno SF, Loughry WJ, editors. The biology of the Xenarthra. Gainesville: University Press of Florida; p. 39–55.
   Google Scholar
• Melchor RN, Perez M, Cardonatto MC, Umazano AM. 2015. Late Miocene ground sloth footprints and their paleoenvironment: megatherichnum oportoi revisited. Palaeogeogr Palaeocl. 439:126–143. doi:10.1016/j.palaeo.2015.02.010.
   Web of Science ®Google Scholar
• Melchor, RN, Perez, M, and Umazano, AM. 2013.Continental trace fossils from the Miocene-Pliocene Río Negro Formation (Río Negro and Buenos Aires provinces), Argentina. II Simposio Latinoamericano de Icnología. SLIC Post-symposium Fieldtrip Guidebook (Universidad Nacional de La Pampa: INCITAP); pp. 31-45.
   Google Scholar
• Miño-Boilini ÁR. 2016. Additions to the knowledge of the ground sloth Catonyx tarijensis (Xenarthra, Pilosa) in the Pleistocene of Argentina. Palaeont Z. 90(1):173–183. doi:10.1007/s12542-015-0280-6.
   Google Scholar
• Miño-Boilini ÁR, Carlini AA. 2009. The Scelidotheriinae Ameghino, 1904 (Phyllophaga, Xenarthra) from the Ensenadan–Lujanian Stage/Ages (Early Pleistocene to Early-Middle Pleistocene–Early Holocene) of Argentina. Quatern Int. 210(1–2):93–101. doi:10.1016/j.quaint.2009.06.016.
   Google Scholar
• Oliva C, Arregui M. 2018. Mammalian Ichnopathology: a case study of Holartic Ungulates (Gomphotheriidae, Equidae, Camelidae) of the Late Pleistocene of South America. Ichnotaxomic implications. Boletín de la Sociedad Geológica Mexicana. 70(2):417–447. doi:10.18268/BSGM2018v70n2a10.
   Web of Science ®Google Scholar
• Oliva C, Arregui M, Lirusso V, de Valais S. 2013. Laguna del Monte, un nuevo yacimiento Paleoicnológico del Pleistoceno tardío (Piso/Edad Lujanense) Guaminí, provincia de Buenos Aires (Argentina). Paper presented at: Second Latin American Symposium on Ichnology (SLIC2013), Abstract and Intrasymposium Fieldtrip Guide 52; Santa Rosa, La Pampa, Argentina.
   Google Scholar
• Orr CM. 2005. Knuckle‐walking anteater: a convergence test of adaptation for purported knuckle-walking features of African Hominidae. Am J Phys Anthropol. 128(3):639–658. doi:10.1002/ajpa.20192.
   PubMed Web of Science ®Google Scholar
• Owen R. 1842. Description of the skeleton of an extinct gigantic sloth, Mylodon robustus owen, with observations on the osteology, natural affinities, and probable habits of the megatherioid quadrupeds in general. London: R&J Taylor.
   Google Scholar
• Owen R. 1859. On the Megatherium (Megatherium americanum, Cuvier and Blumenbach). Part V. Bones of the posterior extremities. Philos T Roy Soc B. 149:809–829.
   Google Scholar
• Polly PD. 2007. Limbs in mammalian evolution. In: Hall BK, editor. Fins into Limbs: evolution, Development, and Transformation. Chicago: University of Chicago Press; p. 245–268.
   Google Scholar
• Scott WB. 1903-1904. Mammalia of the Santa Cruz Beds. I Edentata. Reports of the Princeton University Expeditions to Patagonia. 5:1–226.
   Google Scholar
• Sorkin B. 2006. Ecomorphology of the giant short-faced bears Agriotherium and Arctodus. Hist Biol. 18(1):1–20. doi:10.1080/08912960500476366.
   Google Scholar
• Stevens KA, Ernst S, Marty D. 2016. Uncertainty and ambiguity in the interpretation of sauropod trackways. In: Marty D, Falkingham PL, Richter A, editors. Dinosaur Tracks: the Next Steps. Bloomington: Indiana University Press; p. 227–243.
   Google Scholar
• Stock C. 1917. Structure of the pes in Mylodon harlani. Bulletin of the Deparment of Geology of the University of California. 10(16):267–286.
   Google Scholar
• Stock C. 1920. Origin of the supposed human footprints of Carson City, Nevada. Science. 51(1325):514. doi:10.1126/science.51.1325.514.a.
   PubMedGoogle Scholar
• Stock C. 1925. Cenozoic gravigrade edentates of Western North America: with special reference to the Pleistocene Megalonychinae and Mylodontidae of Rancho La Brea. Washington: Carnegie Institution.
   Google Scholar
• Tassara D, Aramayo S, Osterrieth M, Scian R. 2007. Paleoicnología de mamíferos de la Formación Santa Clara (Pleistoceno tardío) en la zona costera del Partido de Mar Chiquita (Provincia de Buenos Aires) Argentina. Paper presented at: Actas del XVI Congreso Geológico Argentino. 4:219–226.
   Google Scholar
• Thulborn T. 1990. Dinosaur Tracks. London: Chapman and Hall.
   Google Scholar
• Toledo N, De Iuliis G, Vizcaíno SF, Bargo MS. 2018. The concept of a pedolateral pes revisited: the giant sloths Megatherium and Eremotherium (Xenarthra, Folivora, Megatheriinae) as a case study. J Mamm Evol. 25(4):525–537. doi:10.1007/s10914-017-9410-0.
   Web of Science ®Google Scholar
• Vizcaíno SF, Cassini GH, Toledo N, Bargo MS. 2012. On the Evolution of Large Size in Mammalian Herbivores of Cenozoic Faunas of Southern South America. In: In: Patterson BD, Costa LP, editors. Bones, Clones, andBiomes. Chicago: University of Chicago Press; p. 76–101.
   Google Scholar
• Vizcaíno SF, Zárate M, Bargo MS, Dondas A. 2001. Pleistocene burrows in the Mar del Plata area (Argentina) and their probable builders. Acta Palaeontol Pol. 46(2):289–301.
   Web of Science ®Google Scholar
• Weissengruber GE, Egger GF, Hutchinson JR, Groenewald HB, Elsässer L, Famini D, Forstenpointner G. 2006. The structure of the cushions in the feet of African elephants (Loxodonta africana). J Anat. 209(6):781–792. doi:10.1111/j.1469-7580.2006.00648.x.
   PubMed Web of Science ®Google Scholar
• Winge H. 1915. Jordfundne og nulevende Gumlere (Edentata) fra. Lagoa Santa (Minas Geraes, Brasilien. Copenhagen): Bianco Lunos Bookprinting.
   Google Scholar
• Yalden DW. 1966. A contribution to the functional morphology of the mammalian carpus. J Zool. 149(1):55–64. doi:10.1111/j.1469-7998.1966.tb02983.x.
   Google Scholar