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Research Article

Vertebrate ichnology and palaeoenvironmental associations of Alaska’s largest known dinosaur tracksite in the Cretaceous Cantwell Formation (Maastrichtian) of Denali National Park and Preserve

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Received 27 Nov 2022, Accepted 31 May 2023, Published online: 27 Jul 2023

References

  • Blair TC, McPherson JG. 1994. Alluvial fans and their natural distinction from rivers based on morphology, hydraulic processes, sedimentary processes, and facies assemblages. J Sediment Res. A64(3):450–489.
  • Brouwers EM, Clemens WA, Spicer RA, Ager TA, Carter LD, Slitter WV. 1987. Dinosaurs on the North Slope, Alaska: high latitude, Latest Cretaceous environments. Science. 237(4822):1608–1610. doi:10.1126/science.237.4822.1608.
  • Brown CM, Druckenmiller P. 2011. Basal ornithopod (Dinosauria: Ornithischia) teeth from the Prince Creek Formation (Early Maastrichtian) of Alaska. Can J Earth Sci. 48(9):1342–1354. doi:10.1139/e11-017.
  • Chinsamy A, Thomas DB, Tumarkin-Deratzian AR, Fiorillo AR. 2012. Hadrosaurs were perennial polar residents. Anat Rec. 295(4):610–614. doi:10.1002/ar.22428.
  • Cohen KM, Finney SC, Gibbard PL, Fan J-X. 2013. The ICS International Chronostratigraphic Chart. Episodes. 36(3):199–204. doi:10.18814/epiiugs/2013/v36i3/002.
  • Conrad JE, McKee EH, Turrin BD. 1990. Age of tephra beds at the Ocean Point dinosaur locality, North Slope, Alaska, based on K-Ar and 40Ar/39Ar analyses. United States Geological Survey Bulletin. 1990-C:1–12.
  • Coria RA, Currie PJ, Eberth D, Garrido A. 2002. Bird footprints From the Anacleto Formation (Late Cretaceous) in Neuquén Province, Argentina. Ameghiniana. 39(4):1–11.
  • Currie PJ. 1981. Bird footprints from the Gething Formation (Aptian, Lower Cretaceous) of northeastern British Columbia, Canada. J Vertebr Paleontol. 1(3–4):257–264. doi:10.1080/02724634.1981.10011900.
  • Currie PJ, Badamgarav D, Koppelhus E. 2003. The first Late Cretaceous footprints from the Nemegt Locality in the Gobi of Mongolia. Ichnos. 10(1):1–13. doi:10.1080/10420940390235071.
  • Davies KL. 1987. Duck-bill dinosaurs (Hadrosauridae, Ornithischia) from the North Slope of Alaska. J Paleontol. 61(1):198–200. doi:10.1017/S0022336000028341.
  • Díaz-Martínez I, Pereda-Suberbiola X, Pérez-Lorente F, Canudo JI. 2015. Ichnotaxonomic review of large ornithopod dinosaur tracks: temporal and geographic implications. PLoS ONE. 10(2):e0115477. doi:10.1371/journal.pone.0115477.
  • Druckenmiller PS, Erickson GM, Brinkman D, Brown CM, Mori H. 2013. Evidence for a distinct, Early Maastrichtrian polar dinosaur fauna from the Prince Creek Formation of northern Alaska. J Vertebr Paleontol, Programs and Abstracts. 2013:117.
  • Druckenmiller PS, Erickson GM, Benowitz JA, Capps D, Knight C, May KC, Stewart DG. 2017. Deinonychosaurian trace fossils from the Late Cretaceous (Maastrichtian) Cantwell Formation, Alaska: track maker inferences using coeval interformational comparisons. Geol. Soc of Am Abstracts with Programs. 49(6):0016–7592.
  • Druckenmiller PS, Erickson GM, Brinkman D, Brown CM, Eberle JJ. 2021. Nesting at extreme polar latitudes by non-avian dinosaurs. Curr Biol. 31(16):3469–3478. doi:10.1016/j.cub.2021.05.041.
  • Enriquez NJ, Campione NE, White MA, Fanti F, Sissons RL, Sullivan C, Vavrek MJ, Bell PR. 2022. The dinosaur tracks of Tyrants Aisle: an Upper Cretaceous ichnofauna from Unit 4 of the Wapiti Formation (Upper Campanian). PLoS ONE. 17(2):e0262824.
  • Farlow JO. 2018. Noah’s Ravens: interpreting the makers of tridactyl dinosaur footprints. Bloomington (Indiana): Indiana University Press.
  • Fiorillo AR. 2008. On the occurrence of exceptionally large teeth of Troodon (Dinosauria: Saurischia) from the Late Cretaceous of Northern Alaska. PALAIOS. 23(5):322–328. doi:10.2110/palo.2007.p07-036r.
  • Fiorillo AR. 2018. Arctic Dinosaurs: an ancient Arctic world. Boca Raton: CRC Press.
  • Fiorillo AR, Adams TL. 2012. A therizinosaur track from the Lower Cantwell Formation (Upper Cretaceous) of Denali National Park, Alaska. Palaios. 27(6):395–400. doi:10.2110/palo.2011.p11-083r.
  • Fiorillo AR, Contessi M, Kobayashi Y, McCarthy PJ. 2014a. Theropod tracks from the Lower Cantwell Formation (Upper Cretaceous) of Denali National Park, Alaska, USA with comments on theropod diversity in an ancient, high-latitude terrestrial ecosystem. In: Lockley MG, Lucas SG, editors. Fossil footprints of western North America, New Mexico Museum of Natural History and Science Bulletin. 62:429–440.
  • Fiorillo AR, Gangloff RA. 2000. Theropod teeth from the Prince Creek Formation (Cretaceous) of Northern Alaska, with speculations on Arctic dinosaur paleoecology. J Vertebr Paleontol. 20(4):675–682. doi:10.1671/0272-4634(2000)020[0675:TTFTPC]2.0.CO;2.
  • Fiorillo AR, Gangloff RA. 2001. The caribou migration model for Arctic hadrosaurs (Ornithischia: Dinosauria): a reassessment. Hist Biol. 15(4):323–334. doi:10.1080/0891296021000037327.
  • Fiorillo AR, Hasiotis ST, Kobayashi Y. 2014b. Herd structure in Late Cretaceous polar dinosaurs: a remarkable new dinosaur tracksite, Denali National Park: Alaska (USA). Geology. 42(8): 719–722.
  • Fiorillo AR, Hasiotis ST, Kobayashi Y, Breithaupt BH, McCarthy PJ. 2011. Bird tracks from the Upper Cretaceous Cantwell Formation of Denali National Park, Alaska, USA: a new perspective on ancient northern polar vertebrate biodiversity. J Syst Palaeontol. 9(1):33–49. doi:10.1080/14772019.2010.509356.
  • Fiorillo AR, Hasiotis ST, Kobayashi Y, Tomsich CS. 2009b. A pterosaur manus track from Denali National Park, Alaska Range, Alaska, United States. PALAIOS. 24:466–472.
  • Fiorillo AR, Kobayashi Y, McCarthy PJ, Wright TC, Tomsich CS. 2015. Pterosaur tracks from the Lower Cantwell Formation (Campanian–Maastrichtian) of Denali National Park, Alaska, USA, with comments about landscape heterogeneity and habit preferences. Hist Biol. 27(6):672–683. doi:10.1080/08912963.2014.933213.
  • Fiorillo AR, McCarthy PJ, Breithaupt BH, Brease P. 2007. Dinosauria and fossil Aves footprints from the Lower Cantwell Formation (latest Cretaceous), Denali Park and Preserve, Alaska. Alaska Park Service. 6:41–43.
  • Fiorillo AR, McCarthy PJ, Flaig PP. 2010. Taphonomic and sedimentologic interpretations of the dinosaur-bearing Upper Cretaceous Strata of the Prince Creek Formation, Northern Alaska: insights from an ancient high-latitude terrestrial ecosystem. Palaeogeogr Palaeoclimatol Palaeoecol. 295(3–4):376–388. doi:10.1016/j.palaeo.2010.02.029.
  • Fiorillo AR, McCarthy PJ, Kobayashi Y, Tomsich CS, Tykoski RS, Lee Y, Tanaka T, Noto CR. 2018. An unusual association of hadrosaur and therizinosaur tracks within Late Cretaceous rocks of Denali National Park, Alaska. Sci Rep. 8(1):11706. doi:10.1038/s41598-018-30110-8.
  • Fiorillo AR, Tykoski RS. 2012. A new Maastrichtian species of the centrosaurine ceratopsid Pachyrhinosaurus from the North Slope of Alaska. Acta Palaeontol Pol. 57(3):561–573. doi:10.4202/app.2011.0033.
  • Fiorillo AR, Tykoski RS. 2014. A diminutive new tyrannosaur from the top of the world. PloS ONE. 9(3):e91287. doi:10.1371/journal.pone.0091287.
  • Fiorillo AR, Tykoski RS. 2016. Small hadrosaur manus and pes tracks from the Lower Cantwell Formation (Upper Cretaceous) Denali National Park, Alaska: implications for locomotion in juvenile hadrosaurs. PALAIOS. 31(10):479–482. doi:10.2110/palo.2016.049.
  • Fiorillo AR, Tykoski RS, Currie PJ, McCarthy PJ, Flaig PP. 2009a. Description of two partial Troodon braincases from the Prince Creek Formation (Upper Cretaceous), North Slope Alaska. J Vertebr Paleontol. 29(1):178–187. doi:10.1080/02724634.2009.10010370
  • Flaig PP, 2010. Depositional environments of the Late Cretaceous (Maastrichtian) dinosaur-bearing Prince Creek Formation: Colville River Region, North Slope, Alaska (Unpubl. PhD thesis). University of Alaska Fairbanks.
  • Flaig PP, McCarthy PJ, Fiorillo AR. 2013. Anatomy, evolution, and paleoenvironmental interpretation of an ancient Arctic coastal plain: integrated paleopedology and palynology from the Upper Cretaceous (Maastrichtian) Prince Creek Formation, North Slope. Alaska (USA). Society for Sedimentary Geology. p.179–230. 978-1-56576-322-7.
  • Gangloff RA. 1995. Edmontonia sp., The First Record of an ankylosaur from Alaska. J Vertebr Paleontol. 15(1):195–200. doi:10.1080/02724634.1995.10011218
  • Gangloff RA, Fiorillo AR, Norton DW. 2005. The first pachycephalosaurine (Dinosauria) from the paleo-Arctic of Alaska and its paleogeographic implications. J Paleontol. 79(5):997–1001. doi:10.1666/0022-3360(2005)079[0997:TFPDFT]2.0.CO;2
  • Gilbert WG, Ferrell VM, Turner DL. 1976. The Teklanika Formation – a new Paleocene volcanic formation in the Central Alaska Range. Geologic Report. 47:1–16.
  • Golovneva LB. 2000. Aquatic plant communities at the Cretaceous-Paleogene boundary in North-Eastern Russia. Acta Palaeobotanica. 40(2):139–151.
  • Herman AB. 2007. Comparative paleofloristics of the Albian-Early Paleocene in the Anadyr-Koryak and North Alaska subregions, part 2: the North Alaska subregion. Stratigr Geol Correl. 15(4):373–384.
  • Herman AB, Spicer RA, Spicer TEV. 2016. Environmental constraints on terrestrial vertebrate behaviour and reproduction in the high Arctic of the Late Cretaceous. Palaeogeogr Palaeoclimatol Palaeoecol. 441:317–338. doi:10.1016/j.palaeo.2015.09.041.
  • Hillhouse JW, Coe RS. 1994 Paleomagnetic data from Alaska. In: Plafker G, Berg H, editors. The Geology of Alaska: Boulder, Colorado, Geological Society of America, The Geology of North America. G-1:797–812.
  • Hitchcock EH, 1845. An attempt to name, classify, and describe the animals that made the fossil footmarks of New England. Proceedings of the 6th Annual Meeting of the Association of American Geologists and Naturalists, New Haven, Connecticut 6, 23–25.
  • Horstwood MS, Košler J, Gehrels G, Jackson SE, McLean NM, Paton C, Pearson NJ, Sircombe K, Sylvester P, Vermeesch P, et al. 2016. Community‐derived standards for LA‐ICP‐MS U‐(Th‐) Pb geochronology–Uncertainty propagation, age interpretation and data reporting. Geostand Geoanal Res. 40(3):311–332.
  • Hoskin PWO, Schaltegger U. 2003. The composition of zircon and igneous and metamorphic petrogenesis. Rev Mineral Geochem. 53(1):27–62.
  • Huh M, Lockley MG, Kim KS, Kim JY, Gwak S. 2012. First reports of Aquatilavipes from Korea: new finds from Cretaceous strata in the Yeosu Islands Archipelago. Inchnos. 19(1–2):43–49.
  • Kim JY, Kim KS, Lockley MG, Yang SY, Seo SJ, Choi HI, Lim JD. 2008. New didactyl dinosaur footprints (Dromaeosauripus hamanensis ichnogen. et ichnosp. nov.) from the Early Cretaceous Haman Formation, south coast of Korea. Palaeogeogr Palaeoclimatol Palaeoecol. 262(1–2):72–78. doi:10.1016/j.palaeo.2008.02.003
  • Li R, Lockley MG, Makovichy PJ, Matsukawa M, Norell MA, Harris JD, Liu M. 2007. Behavioral and faunal implications of Early Cretaceous deinonychosaur trackways from China. Naturwissenschaften. 95:185–191.
  • Lockley M. 1991. Tracking Dinosaurs. Cambridge (New York, Port Chester, Melbourne, Sydney): Cambridge University Press.
  • Lockley MG, Cart K, Martin J, Milner ARC. 2011a. New theropod tracksites from the Upper Cretaceous “Mesaverde” Group, western Colorado, implications for ornithomimosaur track morphology Fossil Record 3. In: Sullivan R, Lucas S, and Spielmann J, editors. New Mexico Museum of Natural History and Science, Bulletin. Vol. 53; pp. 330–336.
  • Lockley MG, Hunt AP. 1995. Ceratopsid tracks and associated ichnofauna from the Laramie Formation (Upper Cretaceous: Maastrichtian) of Colorado. J Vertebr Paleontol. 15(3):592–614. doi:10.1080/02724634.1995.10011251
  • Lockley MG, Janke PR, Triebold M. 2011b. Tracking Tyrannosaurus: notes on purported T. rex tracks. Ichnos. 18:172–175.
  • Lockley MG, Meyer CA, Dos Santos VF. 1998. Megalosauripus and the problematic concept of megalosaur footprints. Gaia. 15:313–337.
  • Lockley MG, Nadon G, Currie PJ. 2003. A diverse dinosaur-bird footprint assemblage from the Lance Formation, Upper Cretaceous, eastern Wyoming: Implications for Ichnotaxonomy. Ichnos. 11(3–4): 229–249. doi: 10.1080/10420940490428625
  • Lockley MG, Wright JL, Matsukawa M. 2001. A new look at Magnoavipes and so-called “Big Bird” tracks from Dinosaur Ridge (Cretaceous, Colorado). Rocky Mt Geol Assoc Geol. 38(3):137–146.
  • Lockley MG, Xing L, Lockwood JAF, Pond S. 2014. A review of large Cretaceous ornithopod tracks, with special reference to their ichnotaxonomy. Biological Journal of the Linnean Society. 113:721–736.
  • Lucas SG, Klein H, Lockley MG, Spielmann JA, Gierlinksi GD, Hunt AP, Tanner L. 2006. Triassic-Jurassic stratigraphic distribution of the theropod footprint ichnogenus Eubrontes The terrestrial Triassic-Jurassic transition. In: Harris J, Lucas S, Spielmann J, Lockley M, Milner A, and Kirkland J, editors. New Mexico Museum of Natural History and Science Bulletin. Vol. 37; pp. 86–93.
  • Ludwig KR. 2008. User’s Manual for Isoplot 3.70. A geochronologic toolkit for Microsoft Excel. Berkeley Geochronology Center Special Publication. 4:73.
  • Lull RS. 1953. ed. Triassic life of the Connecticut Valley (Rev. Hartford (Connecticut): State of Connecticut Bulletin, State Geological and Natural History Survey 81.
  • McCrea RT, Buckley LG, Farlow JO, Lockley MG, Currie PJ, Matthews NA, Pemberton SG. 2014a. A ‘Terror of Tyrannosaurs’: the first trackways of tyrannosaurids and evidence of gregariousness and pathology in Tyrannosauridae. PLoS ONE. 10(1):e0117606.
  • McCrea RT, Buckley LG, Plint AG, Currie PJ, Haggart JW, Helm CW, Pemberton SG 2014b. A review of vertebrate track-bearing Formations from the Mesozoic and Earliest Cenozoic of western Canada with a description of a new theropod ichnospecies and reassignment of an avian ichnogenus Fossil footprints of western North America. In: Lockley M, and Lucas S, editors. New Mexico Museum of Natural History BulletinVol. 62; pp. 5–93.
  • McCrea RT, Lockley MG, Meyer CA. 2001. Global distribution of purported ankylosaur track occurrences (Chapter 20). In: Carpenter K, editor. The armored dinosaurs. Bloomington and Indianapolis: Indiana University Press; p. 413–454.
  • McCrea RT, Sarjeant WAS. 2001. New ichnotaxa of bird and mammal footprints from the Lower Cretaceous (Albian) Gates Formation of Alberta. In: Tanke DH, Carpenter K, editors. Mesozoic vertebrate life. Bloomington and Indianapolis: Indiana University Press; p. 453–478.
  • Miall AD. 2006. The geology of fluvial deposits. Berlin (Heidelberg): Springer Berlin Heidelberg. doi:10.1007/978-3-662-03237-4
  • Milàn J, Bromley RG. 2006. True tracks, undertracks and eroded tracks, experimental work with tetrapod tracks in laboratory and field. Palaeogeogr Palaeoclimatol Palaeoecol. 231(3–4):253–264. doi:10.1016/j.palaeo.2004.12.022
  • Moratalla JJ, Sanz JL, Jimenez S, Lockley MG. 1992. A quadrupedal ornithopod trackway from the Lower Cretaceous of La Rioja (Spain): inferences on gait and hand structure. J Vertebr Paleontol. 12(2):150–157. doi:10.1080/02724634.1992.10011445
  • Mori H, Druckenmiller PS, Erickson GM. 2014. A new Arctic hadrosaurid from the Prince Creek Formation (Lower Maastrichtian) of Northern Alaska. Acta Palaeontol Pol. 61(1):15–32.
  • Nakajima J, Kobayashi Y, Chinzorig T, Tanaka T, Takasaki R, Tsogtbaatar K, Currie PJ, Fiorillo AR. 2018. Dinosaur tracks at the Nemegt locality: paleobiological and paleoenvironmental implications. Palaeogeogr Palaeoclimatol Palaeoecol. 494:147–159. doi:10.1016/j.palaeo.2017.10.026.
  • Parrish JM, Parrish JT, Hutchinson JH, Spicer RA. 1987. Late Cretaceous vertebrate fossils from the North Slope of Alaska and implications for dinosaur ecology. PALAIOS. 2(4):377–389.
  • Phillips RL. 2003. Depositional environments and processes in Upper Cretaceous nonmarine and marine sediments, Ocean Point dinosaur locality, North Slope, Alaska. Cretac Res. 24:499–523. 5. doi:10.1016/S0195-6671(03)00068-5
  • Renne PR, Deino AL, Walter RC, Turrin BD, Swisher CC III, Becker TA, Curtis GH, Sharp WD, Jaouni AR. 1994. Intercalibration of astronomical and radioisotopic time. Geology. 22:783–786. 9. doi:10.1130/0091-7613(1994)022<0783:IOAART>2.3.CO;2
  • Ridgway KD, Decelles PG. 1993. Stream-dominated alluvial fan and lacustrine depositional systems in Cenozoic strike-slip basins, Denali fault system, Yukon Territory, Canada. Sedimentology. 40:645–666. doi:10.1111/j.1365-3091.1993.tb01354.x
  • Ridgway KD, Trop JM, Nokleberg WJ, Davidson CM, Eastham KR. 2002. Mesozoic and Cenozoic tectonics of the eastern and central Alaska Range: progressive basin development and deformation in a suture zone. GSA Bull. 114(12):1480–1503.
  • Ridgway KD, Trop JM, Sweet AR. 1997. Thrust-top basin formation along a suture zone, Cantwell basin, Alaska Range: implications for development of the Denali fault system. GSA Bull. 109(5):505–523.
  • Salazar-Jaramillo S, 2014. Paleoclimate and paleoenvironment of the Prince Creek and Cantwell Formations, Alaska: Terrestrial evidence of Middle Maastrichtian greenhouse event (Unpubl. PhD thesis). University of Alaska Fairbanks, 1–183.
  • Salazar-Jaramillo S, Fowell SJ, McCarthy PJ, Benowitz JA, Śliwiński MG, Tomsich CS. 2016. Terrestrial isotopic evidence for a Middle-Maastrichtian warming event from the lower Cantwell Formation, Alaska. Palaeogeogr Palaeoclimatol Palaeoecol. 441:360–376. doi:10.1016/j.palaeo.2015.09.044.
  • Smith ND, Cross TA, Dufficy JP, Clough SR. 1989. Anatomy of an avulsion. Sedimentology. 36:1–23. doi:10.1111/j.1365-3091.1989.tb00817.x
  • Spicer RA, Herman AB. 2010. The Late Cretaceous environment of the Arctic: a quantitative reassessment based on plant fossils. Palaeogeogr Palaeoclimatol Palaeoecol. 295(3–4):423–442. 10.1016/j.palaeo.2010.02.025
  • Spicer RA, Parrish JT. 1987. Plant megafossils, vertebrate remains, and paleoclimate of the Kogosukruk Tongue (Late Cretaceous), North Slope, Alaska. U.S. Geol Surv Circ. 998:47–48.
  • Sternberg C. 1926. Dinosaur tracks from the Edmonton Formation of Alberta. Geol Surv Can Bull. 44:85–87.
  • Takasaki R, Fiorillo AR, Kobayashi Y, Tykoski RS, McCarthy PJ. 2019. The first definite lambeosaurine bone from the Liscomb Bonebed of the Upper Cretaceous Prince Creek Formation, Alaska, United States. Sci Rep. 9:5384. doi:10.1038/s41598-019-41325-8
  • Tomsich CS, McCarthy PJ, Fowell SJ, Suderlin D. 2010. Paleofloristic and paleoenvironmental information from the Late Cretaceous (Maastrichtian) flora of the lower Cantwell Formation near Sable Mountain, Denali National Park, Alaska. Palaeogeogr Palaeoclimatol Palaeoecol. 295:389–408.
  • Tomsich CS, McCarthy PJ, Fiorillo AR, Stone DB, Benowitz JA, O’Sullivan PB, 2014. New zircon U-Pb ages for the lower Cantwell Formation: implications for the Late Cretaceous paleoecology and paleoenvironment of the lower Cantwell Formation near Sable Mountain, Denali National Park and Preserve, central Alaska Range, USA. ICAM VI: Proceedings. St. Petersburg: VSEGEI, 19–60.
  • Trop JM, Benowitz J, Cole RB, O’Sullivan P. 2019. Cretaceous to Miocene magmatism, sedimentation, and exhumation within the Alaska Range suture zone: a polyphase reactivated terrane boundary. Geosphere. 15(4):1066–1101. doi:10.1130/GES02014.1
  • Thulborn T. 1990. Dinosaur Tracks. London, New York, Tokyo, Melbourne, Madras: Chapman & Hall.
  • Vialov OS. 1988. On the classification of dinosaurian traces [in Russian]. Ežegodnik Vsesoûznogo Paleontologičeskogo Obŝestva. 31:322–325.
  • Walker JD, Geissman JW, Compilers. 2022. Geologic Time Scale v. 6.0: geological Society of America. doi:10.1130/2022.CTS006C.
  • Watanabe A, Erickson GM, Druckenmiller PS. 2013. An ornithomimosaurian from the Upper Cretaceous Prince Creek Formation of Alaska. J Vertebr Paleontol. 33(5):1169–1175. doi:10.1080/02724634.2013.770750
  • Wolfe JA, Wahrhaftig C. 1968. The Cantwell Formation of the Central Alaska Range. Cohee GV, Bates RG, Wright WB editors Changes in stratigraphic nomenclature by the U.S. Geological Survey. 1968 Contributions to Stratigraphy, Geol Surv Bull 1294-A, A41–A55.
  • Xing LD, Harris JD, Sun DH, Zhao HQ. 2009. The earliest known deinonychosaur tracks from the Jurassic–Cretaceous boundary in Hebei, China. Acta Palaeontologica Sinica. 48:662–671.
  • Xing L, Niu K, Lockley MG, Klein H, Romilio A, Persons IV, W.S, Brusatte SL. 2019. A probable tyrannosaurid track from the Upper Cretaceous of southern China. Sci Bull. 64(16):1136–1139.

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