Revisiting the population of the Ghost Crab, Ocypode cursor, on the sandy beaches of northern Cyprus after two decades: are there causes for concern?

References

• Aydın, M. (2011). Growth, reproduction and diet of Pufferfish (Lagocephalus sceleratus Gmelin, 1789) from Turkey’s Mediterranean Sea Coast. Turkish Journal of Fisheries and Aquatic Sciences, 11, 569–576. doi: 10.4194/trjfas.2011.0318
   Web of Science ®Google Scholar
• Broderick, A. C., & Godley, B. J. (1996). Population and nesting ecology of the Green Turtle, Chelonia mydas, and the Loggerhead Turtle, Caretta caretta, in northern Cyprus. Zoology in the Middle East, 13, 27–46. doi: 10.1080/09397140.1996.10637704
   Google Scholar
• Carbonara, P., Kolitari, J., Djurovic, M., Gaudio, P., Ikica, Z., Kroqi, G., Milone, N., Spedicato, M. T. (2017). The presence of Tetraodontidae species in the Central Mediterranean: an update from the southern Adriatic Sea. Acta Adriatica, 58, 325–338. doi: 10.32582/aa.58.2.11
   Web of Science ®Google Scholar
• Hereward, H. F. R., Gentle, L. K., Ray, N. D., & Sluka, R. D. (2017). Ghost Crab burrow density at Watamu Marine National Park: An indicator of the impact of urbanisation and associated disturbance? African Journal of Marine Science, 39, 129–133. doi: 10.2989/1814232X.2017.1305990
   Web of Science ®Google Scholar
• Kasparek, M., Godley, B. J., & Broderick, A. C. (2001). Nesting of the Green Turtle, Chelonia mydas, in the Mediterranean: a review of status and conservation needs. Zoology in the Middle East, 24, 45–74. doi: 10.1080/09397140.2001.10637885
   Google Scholar
• Jonah, F. E., Aheto, D. W., Adjei-Boateng, D., Agbo, N. W., Boateng, I., & Shimba, M. J. (2015). Human use and modification of beaches and dunes are linked to Ghost Crab (Ocypode spp) population decline in Ghana. Regional Studies in Marine Science, 2, 87–94. doi: 10.1016/j.rsma.2015.08.013
   Web of Science ®Google Scholar
• Lucrezi, S. (2015). Ghost Crab populations respond to changing morphodynamic and habitat properties on sandy beaches. Acta Oecologica, 62, 18–31. doi: 10.1016/j.actao.2014.11.004
   Web of Science ®Google Scholar
• Lucrezi, S., Saayman, M., & van der Merwe, P. (2014). Impact of off-road vehicles (ORVs) on ghost crabs of sandy beaches with traffic restrictions: A case study of Sodwana Bay, South Africa. Environmental Management, 53, 520–533. doi: 10.1007/s00267-013-0223-5
   PubMed Web of Science ®Google Scholar
• Marco, A., da Graça, J., García-Cerdá, R., Abella, E., & Freitas, R. (2015). Patterns and intensity of Ghost Crab predation on the nests of an important endangered loggerhead turtle population. Journal of Experimental Marine Biology and Ecology, 468, 74–82. doi: 10.1016/j.jembe.2015.03.010
   Web of Science ®Google Scholar
• Oliveira, C. A. G. de, Souza, G. N., & Soares-Gomes, A. (2016). Measuring burrows as a feasible non-destructive method for studying the population dynamics of Ghost Crabs. Marine Biodiversity, 46, 809–817. doi: 10.1007/s12526-015-0436-3
   Web of Science ®Google Scholar
• Pombo, M., de Oliveira, A. L., Xavier, L. Y., Siegle, E., & Turra, A. (2017). Natural drivers of distribution of Ghost Crabs Ocypode quadrata and the implications of estimates from burrows. Marine Ecology Progress Series, 565, 131–147. doi: 10.3354/meps11991
   Web of Science ®Google Scholar
• Sakai, K., & Türkay, M. (2013). Revision of the genus Ocypode with the description of a new genus, Hoplocypode (Crustacea: Decapoda: Brachyura). Memoirs of the Queensland Museum– Nature, 56, 665–793.
   Google Scholar
• Schlacher, T. A., de Jager, R., & Nielsen, T. (2011). Vegetation and Ghost Crabs in coastal dunes as indicators of putative stressors from tourism. Ecological Indicators, 11, 284–294. doi: 10.1016/j.ecolind.2010.05.006
   Web of Science ®Google Scholar
• Schlacher, T. A., Lucrezi, S., Connolly, R. M., Peterson, C. H., Gilby, B. L., Maslo, B., … Schoeman, D. S. (2016). Human threats to sandy beaches: A meta-analysis of ghost crabs illustrates global anthropogenic impacts. Estuarine, Coastal and Shelf Science, 169, 56–73. doi: 10.1016/j.ecss.2015.11.025
   Web of Science ®Google Scholar
• Steiner, A. J., & Leatherman, S. P. (1981). Recreational impacts on the distribution of Ghost Crabs Ocypode quadrata Fab. Biological Conservation, 20, 111–122. doi: 10.1016/0006-3207(81)90022-7
   Web of Science ®Google Scholar
• Stelling-Wood, T. P., Clark, G. F., & Poore, A. G. B. (2016). Responses of ghost crabs to habitat modification of urban sandy beaches. Marine Environmental Research, 116, 32–40. doi: 10.1016/j.marenvres.2016.02.009
   PubMed Web of Science ®Google Scholar
• Strachan, P. H., Smith, R. C., Hamilton, D. A. B., & Taylor, A. C. (1999). Studies on the ecology and behaviour of the Ghost Crab, Ocypode cursor (L.) in northern Cyprus. Scientia Marina, 63, 51–60. doi: 10.3989/scimar.1999.63n151
   Web of Science ®Google Scholar
• Suciu, M. C., Tavares, D. C., & Zalmon, I. R. (2018). Comparative evaluation of crustaceans as bioindicators of human impact on Brazilian sandy beaches. Journal of Crustacean Biology, 38, 420–428. doi: 10.1093/jcbiol/ruy027
   Web of Science ®Google Scholar
• Rodrigues, E., Freitas, R., Delgado, N. de C., & Soares-Gomes, A. (2016). Distribution patterns of the ghost crab Ocypode cursor on sandy beaches of a tropical island in the Cabo Verde archipelago, Eastern Central Atlantic. African Journal of Marine Science, 38, 181–188 doi: 10.2989/1814232X.2016.1176602
   Web of Science ®Google Scholar
• Shuchman, E., & Warburg, M. R. (1978). Dispersal, population structure and burrow shape of Ocypode cursor. Marine Biology, 49, 255–263. doi: 10.1007/BF00391138
   Web of Science ®Google Scholar
• Türeli, C., Duysak, Ö., Akamca, E., & Kiyagi, V. (2009). Spatial distribution and activity pattern of the Ghost Crab, Ocypode cursor (L., 1758) in Yumurtalik Bay, North-Eastern Mediterranean-Turkey. Journal of Animal and Veterinary Advances, 8, 165–171.
   Google Scholar
• Turra, A., Gonçalves, M. A. O., & Denadai, M. R. (2005). Spatial distribution of the ghost crab Ocypode quadrata in low-energy tide-dominated sandy beaches. Journal of Natural History, 39, 2163–2177. doi: 10.1080/00222930500060165
   Web of Science ®Google Scholar
• Valero-Pacheco, E., Alvarez, F., Abarca-Arenas, L. G., & Escobar, M. (2007). Population density and activity pattern of the Ghost Crab, Ocypode quadrata, in Veracruz, Mexico. Crustaceana, 80, 313–325. doi: 10.1163/156854007780162479
   Web of Science ®Google Scholar
• Wolcott, T. G. (1978). Ecological role of Ghost Crabs, Ocypode quadrata (Fabricius) on an ocean beach: Scavengers or predators? Journal of Experimental Marine Biology and Ecology, 31, 67–82. doi: 10.1016/0022-0981(78)90137-5
   Web of Science ®Google Scholar