Early Triassic Archosaur-Dominated Footprint Assemblage from the Argana Basin (Western High Atlas, Morocco)

REFERENCES

• Ait Chayeb , E , Youbi , N , El-Boukhari , A , Bouabdelli , M and Amrhar , M . 1998 . Le volcanisme permien et mésozoique inférieur du bassin d’Argana (Haut-Atlas occidental, Maroc): un magmatisme intraplaque associé à l’ouverture de l’Atlantique central . Journal of African Earth Sciences , 26 : 499 – 519 .
   Google Scholar
• Avanzini , M. and Mietto , P. 2008 . Lower and Middle Triassic footprint-based Biochronology in the Italian Southern Alps . Oryctos , 8 : 3 – 13 .
   Google Scholar
• Avanzini , M. and Renesto , S. 2002 . A review of Rhynchosauroides tirolicus Abel, 1926 ichnospecies (Middle Triassic: Anisian-Ladinian) and some inferences on Rhynchosauroides trackmaker . Rivista Italiana di Paleontologia e Stratigrafia , 108 ( 1 ) : 51 – 66 .
   Web of Science ®Google Scholar
• Avanzini , M. , Ceoloni , P. , Conti , M. A. , Leonardi , G. , Manni , R. , Mariotti , N. , Mietto , P. , Muraro , C. , Nicosia , U. , Sacchi , E. , Santi , G. and Spezzamonte , M. 2001 . Permian and Triassic tetrapod ichnofaunal units of northern Italy: Their potential contribution to continental biochronology. Natura Bresciana (Annali del Museo Civico di Scienze Naturali di Brescia) . Monografia N , 25 : 89 – 107 .
   Google Scholar
• Bachmann , G. H. and Kozur , H. W. 2004 . The Germanic Triassic: Correlations with the international chronostratigraphic scale, numerical ages and Milankovitch cyclicity . Hallesches Jahrbuch für Geowissenschaften B , 26 : 17 – 62 .
   Google Scholar
• Benton , M. J. 1999 . Scleromochlus taylori and the origin of dinosaurs and pterosaurs . Philosophical Transactions of the Royal Society of London B , 354 : 1423 – 1446 .
   Web of Science ®Google Scholar
• Benton , M. J. 2008 . Presidential address 2007: The end-Permian mass extinction—events on land in Russia . Proceedings of the Geologists’ Association , 119 : 119 – 136 .
   Web of Science ®Google Scholar
• Benton , M. J. , Tverdokhlebov , V. P. and Surkov , M. V. 2004 . Ecosystem remodeling among vertebrates at the Permian-Triassic boundary in Russia . Nature , 432 : 97 – 100 .
   PubMed Web of Science ®Google Scholar
• Biron , P. E. and Dutuit , J.-M. 1981 . Figurations sédimentaires et traces d’activité au sol dans le Trias de la formation d’Argana et de l’Ourika (Maroc) . Bulletin du Muséum National d’ Histoire Naturelle Paris, 4e série, 3, section C , 4 : 399 – 427 .
   Google Scholar
• Botha , J. and Smith , R. M.H. 2006 . Rapid vertebrate recuperation in the Karoo Basin of South Africa following the end-Permian extinction . Journal of African Earth Sciences , 45 : 502 – 514 .
   Web of Science ®Google Scholar
• Brown , R. H. 1980 . Triassic rocks of Argana Valley, Southern Morocco and their regional structural implications . Bulletin of the American Association of Petroleum Geologists , 64 : 988 – 1003 .
   Web of Science ®Google Scholar
• Brusatte , S. L. , Benton , M. J. , Ruta , M. and Lloyd , G. T. 2008 . Superiority, competition, and opportunism in the evolutionary radiation of dinosaurs . Science , 321 : 1485 – 1488 .
   PubMed Web of Science ®Google Scholar
• Conti , M. A. , Leonardi , G. , Mariotti , N. and Nicosia , U. 1977 . Tetrapod footprints of the “Val Gardena Sandstone” (North Italy) . Paleontographica Italiana N.S. , 40 : 91
   Google Scholar
• Deenen , M. H.L. , Ruhl , M. , Bonis , N. R. , Krijgsman , W. , Kuerschner , W. M. , Reitsma , M. and van Bergen , M. J . 2010 . A new chronology for the end-Triassic mass extinction . Earth and Planetary Science Letters , 291 : 113 – 125 .
   Web of Science ®Google Scholar
• Demathieu , G. 1970 . Les empreintes de pas de vertébrés du Trias de la bordure Nord-Est du Massif Central . Cahiers de Paleontologie , : 211
   Google Scholar
• De Koning , G . 1957 . Geologie des Ida ou Zal (Maroc). Stratigraphie, pétrographie et tectonique de la partie SW du bloc occidental du massif ancien du Haut-Atlas . Leidse Geologische Mededelingen , 22 : 209
   Google Scholar
• Demathieu , G. and Demathieu , P. 2004 . Chirotheria and other ichnotaxa of the European Triassic . Ichnos , 11 : 79 – 88 .
   Google Scholar
• Demathieu , G. and Haubold , H. 1972 . Stratigraphische Aussagen der Tetrapodenfährten aus der terrestrischen Trias Europas . Geologie , 21 : 802 – 836 .
   Google Scholar
• Demathieu , G. and Haubold , H. 1982 . Reptilfährten aus dem Mittleren Buntsandstein von Hessen . Hallesches Jahrbuch für Geowissenschaften , 7 : 97 – 110 .
   Google Scholar
• Demathieu , G. and Oosterink , H. W. 1983 . Die Wirbeltier-Ichnofauna aus dem Unteren Muschelkalk von Winterswijk (Die Reptilfährten aus der Mitteltrias der Niederlande) . Staringia , 7 : 1 – 51 .
   Google Scholar
• Dilkes , D. and Sues , H.-D. 2009 . Redescription and phylogenetic relationships of Doswellia kaltenbachi (Diapsida: Archosauriformes) from the Upper Triassic of Virginia . Journal of Vertebrate Paleontology , 29 ( 1 ) : 58 – 79 .
   Web of Science ®Google Scholar
• Dutuit , J.-M. 1972 . Découverte d’un Dinosaure ornithischien dans le Trias supérieur de I’Atlas occidental marocain . Comptes rendus de l′Academie des Sciences , 275 : 2841 – 2844 .
   Google Scholar
• Dutuit , J.-M. 1976a . Introduction à l’étude paléontologique du Trias continental marocain. Description des premiers Stégocéphales recueillis dans le couloir d’Argana (Atlas occidental) . Mémoires du Muséum National d’ Histoire Naturelle, Paris , 36 : 1 – 253 . C
   Google Scholar
• Dutuit , J.-M. 1976b . II est probable que les Rhynchocéphales sont représentés dans la faune du Trias marocain . Comptes Rendus de l’Academie des Sciences, Paris , 238 : 483 – 486 .
   Google Scholar
• Dutuit , J.-M. 1977a . Paleorhinus magnoculus Phytosaure du Trias supérieur de I’Atlas marocain . Annales de l’Université de Provence, Géologie Méditerranéenne , 4 ( 3 ) : 255 – 268 .
   Google Scholar
• Dutuit , J.-M. 1977b . Description du crâne de Angistorhinus talainti n. sp. un nouveau phytosaure du Trias atlasique marocain . Bulletin du Muséum National d’Histoire Naturelle, Paris , 489 : 297 – 336 . Série 3
   Google Scholar
• Dutuit , J.-M. 1979 . Un pseudosuchien du Trias continental marocain . Annales de Paléontologie (Vertébrés) , 65 : 55 – 68 .
   Google Scholar
• Dutuit , J.-M. 1988 . Diplocaulus minimus n. sp. (Amphibia, Nectridea) lépospondyle de la formation d’Argana, dans l’Atlas occidental marocain . Comptes rendus de l’Académie des Sciences, Paris , 307 : 851 – 854 .
   Google Scholar
• Fichter , J. and Lepper , J. 1997 . Die Fährtenplatte vom Heuberg bei Gieselwerder . Philippia , 8/1 : 35 – 60 .
   Google Scholar
• Fichter , J. and Kunz , R. 2004 . New genus and species of chirotheroid tracks in the Detfurth-Formation (Middle Bunter, Lower Triassic) of Central Germany . Ichnos , 11 : 183 – 193 .
   Google Scholar
• Fuglewicz , R. , Ptaszynski , T. and Rdzanek , K. 1990 . Lower Triassic footprints from the Swietokrzyskie (Holy Cross) Mountains, Poland . Acta Palaeontologica Polonica , 35 : 109 – 164 .
   Google Scholar
• Gand , G. 1979 . Note sur quelques nouvelles pistes de reptiles observées dans le Trias moyen du Plateau d’Antully (Saône-et-Loire—France) . Bulletin de la Société d’Histoire Naturelle d’Autun , 92 : 7 – 20 .
   Google Scholar
• Gand , G. and Demathieu , G. 2005 . Les pistes dinosauroides du Trias moyen francais: Interprétation et réévaluation de la nomenclature . Geobios , 38 : 725 – 749 .
   Web of Science ®Google Scholar
• Gauffre , F.-X. 1993 . The prosauropod dinosaur Azendohsaurus laaroussii from the Upper Triassic of Morocco . Palaeontology , 36 ( 4 ) : 897 – 908 .
   Google Scholar
• Ginsburg , L. , de Lapparent , A. F. and Taquet , P. 1968 . Piste de Chirotherium dans le Trias du Niger . Comptes Rendus de l’ Académie des Sciences Paris Série D , 266 : 2056 – 2058 .
   Google Scholar
• Gower , D. J. 1996 . The tarsus of erythrosuchid archosaurs, and implications for early diapsid phylogeny . Zoological Journal of the Linnean Society , 116 : 347 – 375 .
   Web of Science ®Google Scholar
• Gower , D. J. and Schoch , R. R. 2009 . Postcranial anatomy of the rauisuchian archosaur Batrachotomus kupferzellensis . Journal of Vertebrate Paleontology , 29 ( 1 ) : 103 – 122 .
   Web of Science ®Google Scholar
• Gower , D. J. and Sennikov , A. G. 2000 . “ Early archosaurs from Russia ” . In The Age of Dinosaurs in Russia and Mongolia , Edited by: Benton , M. J. , Shishkin , M. A. , Unwin , D. M. and Kurochkin , E. N. 140 – 159 . Cambridge : Cambridge University Press .
   Google Scholar
• Haubold , H. 1966 . Therapsiden- und Rhynchocephalen-Fährten aus dem Buntsandstein Südthüringens . Hercynia , 3 ( 2 ) : 147 – 183 . NF
   Google Scholar
• Haubold , H. 1967 . Eine Pseudosuchier-Fährtenfauna aus dem Buntsandstein Südthüringens . Hallesches Jahrbuch für Mitteldeutsche Erdgeschichte , 8 : 12 – 48 .
   Google Scholar
• Haubold , H. 1969 . Parallelisierung terrestrischer Ablagerungen der tieferen Trias mit Pseudosuchier-Fährten . Geologie , 18 : 836 – 843 .
   Google Scholar
• Haubold , H. 1971a . Die Tetrapodenfährten des Buntsandsteins . Paläontologische Abhandlungen A , : 395 – 548 . IV
   Google Scholar
• Haubold , H. 1971b . Ichnia Amphibiorum et Reptiliorum fossilium . Encyclopedia of Paleoherpetology , 18 : 1 – 124 .
   Google Scholar
• Haubold , H. 1984 . “ Saurierfährten ” . In Wittenberg 231 Ziemsen
   Google Scholar
• Haubold , H. 2006 . Die Saurierfährten Chirotherium barthii Kaup, 1835—das Typusmaterial aus dem Buntsandstein bei Hildburghausen/Thüringen und das Chirotherium-Monument . Veröffentlichungen Naturhistorisches Museum Schleusingen , 21 : 3 – 31 .
   Google Scholar
• Haubold , H. and Klein , H. 2000 . Die dinosauroiden Fährten Parachirotherium–Atreipus–Grallator aus dem unteren Mittelkeuper (Obere Trias: Ladin, Karn, ?Nor) in Franken . Hallesches Jahrbuch für Geowissenschaften B , 22 : 59 – 85 .
   Google Scholar
• Haubold , H. and Klein , H. 2002 . Chirotherien und Grallatoriden aus der Unteren bis Oberen Trias Mitteleuropas und die Entstehung der Dinosauria . Hallesches Jahrbuch für Geowissenschaften B , 24 : 1 – 22 .
   Google Scholar
• Hmich , D. , Schneider , W. , Saber , H. , Voigt , S. and El Wartiti , M. 2006 . New continental Carboniferous and Permian faunas of Morocco: Implications for biostratigraphy, palaeobiogeography and palaeoclimate . Geological Society London, Special Publications , 265 : 297 – 324 .
   Google Scholar
• Hminna , A. , Saber , H. , Schneider , J. W. , Klein , H. , Voigt , S. and Hmich , D. 2009a . Brachychirotherium ichnofauna from the Upper Triassic of Oued Oum Er Rbiaa Formation of Sidi Saïd M’aachou basin, Coastal Meseta, Morocco . 1st International Congress on North African Vertebrate Palaeontology , : 32 Abstracts
   Google Scholar
• Hminna , A. , Voigt , S. , Saber , H. , Schneider , J. W. and Hmich , D. 2009b . Tetrapod footprints from Ikakern Formation (Argana Basin, Western High Atlas, Morocco) . 1st International Congress on North African Vertebrate Palaeontology , : 61 – 62 . Abstracts
   Google Scholar
• Hofmann , A. , Tourani , A. and Gaupp , R. 2000 . Cyclicity of Triassic to Lower Jurassic continental red beds of the Argana Valley, Morocco: Implications for palaeoclimate and basin evolution . Palaeogeography, Palaeoclimatology, Palaeoecology , 161 : 229 – 266 .
   Web of Science ®Google Scholar
• Jalil , N.-E. 1999 . Continental Permian and Triassic vertebrate localities from Algeria and Morocco and their stratigraphical correlations . Journal of African Earth Sciences , 29 ( 1 ) : 219 – 226 .
   Web of Science ®Google Scholar
• Jalil , N.-E. and Dutuit , J.-M. 1996 . Permian captorhinid reptiles from the Argana Formation, Morocco . Palaeontology , 39 ( 4 ) : 907 – 918 .
   Google Scholar
• Jalil , N.-E. and Janvier , P. 2005 . Les paréiasaures (Amniota, Parareptilia) du Permien supérieur du Bassin d’Argana, Maroc . Geodiversitas , 27 : 35 – 132 .
   Google Scholar
• Jalil , N.-E. and Peyer , K. 2007 . A new rauisuchian (Archosauria, Suchia) from the Upper Triassic of the Argana Basin, Morocco . Palaeontology , 50 ( 2 ) : 417 – 430 .
   Web of Science ®Google Scholar
• Jalil , N.-E. , Janvier , P. and Steyer , J.-S. 2009 . A new cyclotosaurid (Amphibia, Temnospondyli) from the Triassic of Argana Basin (High Atlas Mountains, Morocco); biostratigraphic implications . 1st International Congress on North African Vertebrate Palaeontology , : 36 – 37 . Abstracts:
   Google Scholar
• Jones , D. F. 1975 . Stratigraphy, environments of deposition, petrology, age and provenance of the basal red beds of the Argana valley, western High Atlas Mountains, Morocco . New Mexico Institute of Mining and Technology , : 148 Master Thesis
   Google Scholar
• Karl , C. and Haubold , H. 1998 . Brachychirotherium aus dem Coburger Sandstein (Mittlerer Keuper, Karn/Nor) in Nordbayern . Hallesches Jahrbuch für Geowissenschaften B , 20 : 33 – 58 .
   Google Scholar
• Khaldoune , F. , Jalil , N.-E. and Peyer , K. 2009 . Les vertebres fossiles du bassin d’Argana (Permien-Trias, Haut Atlas occidental): Liste faunique et niveaux stratigraphiques . 1st International Congress on North African Vertebrate Palaeontology , : 68 Abstracts
   Google Scholar
• King , M. J. , Sarjeant , W. A.S. , Thompson , D. B. and Tresise , G. 2005 . A revised systematic ichnotaxonomy and review of the vertebrate footprint ichnofamily Chirotheriidae from the British Triassic . Ichnos , 12 : 241 – 299 .
   Google Scholar
• Klein , H. and Haubold , H. 2004 . Überlieferungsbedingte Variation bei Chirotherien und Hinweise zur Ichnotaxonomie nach Beispielen aus der Mittel- bis Ober-Trias (Anisium–Karnium) von Nordbayern . Hallesches Jahrbuch für Geowissenschaften B , 26 : 1 – 15 .
   Google Scholar
• Klein , H. and Haubold , H. 2007 . Archosaur footprints—potential for biochronology of Triassic continental sequences . New Mexico Museum of Natural History and Science, Bulletin , 41 : 120 – 130 .
   Google Scholar
• Klein , H. and Lucas , S. G. 2010 . Tetrapod footprints—their use in biostratigraphy and biochronology of the Triassic . Geological Society, London, Special Publications , 334 : 419 – 446 .
   Google Scholar
• Klein , H. , Saber , H. , Voigt , S. , Schneider , J. W. , Hmich , D. and Hminna , A. 2009 . An archosaur-dominated footprint assemblage in Permo-Triassic red-beds of Morocco and the global record of early chirotherians . 1st International Congress on North African Vertebrate Palaeontology , : 18 Abstracts
   Google Scholar
• Kotanski , Z. , Gierlinski , G. and Ptaszynski , T. 2004 . Reptile tracks (Rotodactylus) from the Middle Triassic of the Djurdjura Mountains in Algeria . Geological Quarterly , 48 ( 1 ) : 89 – 96 .
   Web of Science ®Google Scholar
• Kozur , H. W. and Bachmann , G. H. 2005 . Correlation of the Germanic Triassic with the international scale . Albertiana , 32 : 21 – 35 .
   Google Scholar
• Krebs , B. 1965 . Ticinosuchus ferox n. g. n. sp., ein neuer Pseudosuchier aus der Trias des Monte San Giorgio . Schweizer Paläontologische Abhandlungen , 81 : 1 – 141 .
   Google Scholar
• Kubo , T. and Benton , M. J. 2007 . Evolution of hindlimb posture in archosaurs: Limb stresses in extinct vertebrates . Palaeontology , 50 ( 6 ) : 1519 – 1529 .
   Web of Science ®Google Scholar
• Leonardi , G. 1987 . Glossary and Manual of Tetrapod Footprint Palaeoichnology . Ministerio Minas Energie, Departemento Nacional da Producao Mineral, Brasilia , : 117
   Google Scholar
• Lucas , S. G. 1998 . Global Triassic tetrapod biostratigraphy and biochronology . Palaeogeography, Palaeoclimatology, Palaeoecology , 143 : 347 – 384 .
   Web of Science ®Google Scholar
• Lucas , S. G. 1999 . Tetrapod-based correlation of the nonmarine Triassic . Zentralblatt für Geologie und Paläontologie I , 7–8 : 497 – 521 .
   Google Scholar
• Lucas , S. G. 2003 . Triassic tetrapod footprint biostratigraphy and biochronology . Albertiana , 28 : 75 – 84 .
   Google Scholar
• Lucas , S. G. 2007 . Tetrapod footprint biostratigraphy and biochronology . Ichnos , 14 : 5 – 38 .
   Google Scholar
• MacDonald , D. I.M. , Isbell , J. L. and Hammer , W. R. 1991 . Vertebrate trackways from the Triassic Fremouw Formation, Queen Alexandra Range, Antarctica . Antarctic Journal of the United States , 26 : 20 – 21 .
   Google Scholar
• Maidwell , F. T. 1911 . Notes on footprints from the Keuper of Runcorn Hill . Proceedings of the Liverpool Geological Society , 11 : 140 – 152 .
   Google Scholar
• Medina , F. , Vachard , D. , Colin , J.-P. , Ouarhache , D. D. and Ahmamou , M. 2001 . Charophytes et ostracodes du niveau carbonate de Taourirt Imzilen (Membre d’Aglegal, Trias d’Argana); implications stratigraphiques . Bulletin de l’ Institut scientifique, Rabat, section Sciences de la Terre , 23 : 21 – 26 .
   Google Scholar
• Niedzwiedzki , G. and Ptaszynski , T. 2007 . Large Chirotheriidae tracks in the Early Triassic of Wióry, Holy Cross Mountains, Poland . Acta Geologica Polonica , 57 ( 3 ) : 325 – 342 .
   Web of Science ®Google Scholar
• Nopcsa , F. von. 1923 . Die Familien der reptilien . Fortschritte der Geologie und Paläontologie , 2 : 1 – 210 .
   Google Scholar
• Olsen , P. E. 1997 . Stratigraphic record of the early Mesozoic breakup of Pangea in the Laurasia-Gondwana rift system . Annual Review of Earth and Planetary Science , 25 : 337 – 401 .
   Web of Science ®Google Scholar
• Olsen , P. E. and Et-Touhami , M. 2008 . Fieldtrip Guidebook Central Atlantic Conjugate Margins Conference, Halifax
   Google Scholar
• Olsen , P. E. , Kent , D. V. , Fowell , S. J. , Schlische , R. W. , Withjack , M. O. and LeTourneau , P. M. 2000 . “ Implications of a comparison of the stratigraphy and depositional environments of the Argana (Morocco) and Fundy (Nova Scotia, Canada) Permian-Jurassic basins ” . In Le Permien et le Trias du Maroc , Edited by: Oujidi , M. and Et-Touhami , M. 165 – 183 . Hilal Impression Oujda : Actes de la Première Réunion du Groupe Marocain du Permien et du Trias .
   Google Scholar
• Olsen , P. E. , Kent , D. V. , Et-Touhami , M. , Fowell , S. J. and Witte , B. 2002 . “ Cyclicity, time scale, and correlation of the Triassic and Early Jurassic rift strata of the Argana basin, Morocco ” . In Troisème Réunion du Groupe Marocain du Permien et du Trias , Edited by: Essamoud , R. and Ekamel , F. 29 – 30 . Casablanca . 18–20 Avril, Abstracts
   Google Scholar
• Padian , K. , Li , C. and Pchelnikova , J. 2010 . The trackmaker of Apatopus (Late Triassic, North America): Implications for the evolution of archosaur stance and gait . Palaeontology , 53 ( 1 ) : 175 – 189 .
   Web of Science ®Google Scholar
• Parrish , M. J. 1989 . “ Phylogenetic patterns in the manus and pes of early Mesozoic Archosauromorpha ” . In Dinosaur Tracks and Traces , Edited by: Gillette , D. D. and Lockley , M. G. 249 – 258 . Cambridge University Press : Cambridge, UK .
   Google Scholar
• Peabody , F. E. 1948 . Reptile and amphibian trackways from the Moenkopi Formation of Arizona and Utah . University of California Publications, Bulletin of the Department of Geological Sciences , 27 : 295 – 468 .
   Google Scholar
• Ptaszynski , T. 2000 . Lower Triassic vertebrate footprints from Wióry, Holy Cross Mountains, Poland . Acta Palaeontologica Polonica , 45 : 151 – 194 .
   Web of Science ®Google Scholar
• Retallack , G. J. 1996 . Early Triassic therapsid footprints from the Sydney basin, Australia . Alcheringa , 20 : 301 – 314 .
   Web of Science ®Google Scholar
• Sahney , S. and Benton , M. J. 2008 . Recovery from the most profound mass extinction of all time . Proceedings of the Royal Society B , 275 : 759 – 765 .
   PubMed Web of Science ®Google Scholar
• Schlische , R. W. 2003 . “ Progress in understanding the structural geology, basin evolution, and tectonic history of the eastern North American rift system ” . In The Great Rift Valleys of Pangaea in Eastern North America , Edited by: Le Tourneau , P. M. and Olsen , P. E. 21 – 64 . New York : Vol. 1. Columbia University Press .
   Google Scholar
• Sereno , P. C. 1991 . Basal archosaurs: phylogenetic relationships and functional implications . Journal of Vertebrate Paleontology , 11 ( Suppl. ) : 1 – 65 .
   Google Scholar
• Sereno , P. C. and Arcucci , A. B. 1990 . The monophyly of crurotarsal archosaurs and the origin of bird and crocodile ankle joints . Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen , 180 : 21 – 52 .
   Google Scholar
• Silvestri , S. M. and Szajna , M. J. 1993 . “ Biostratigraphy of vertebrate footprints in the Late Triassic section of the Newark Basin, Pennsylvania: Reassessment of stratigraphic ranges ” . In The Nonmarine Triassic Edited by: Lucas , S. G. and Morales , M. 439 – 444 . New Mexico Museum of Natural History and Science Bulletin, 3
   Google Scholar
• Steyer , J.-S. , Jalil , N.-E. , Taquet , P. , Bitam , I. , Aitouali , R. , Nedjari , A. , Bouzidi , O. and Kedadra , B. 2009 . Temnospondyls from North Africa: Review and perspectives . 1st International Congress on North African Vertebrate Palaeontology , Abstracts: 62
   Google Scholar
• Taquet , P. 1976 . Géologie et Paléontologie du gisement de Gadoufaoua (Aptien du Niger) . Cahiers de Paléontologie, CNRS, Paris , : 1 – 191 .
   Google Scholar
• Tixeront , M. 1973 . Lithostratigraphie et minéralisation cuprifères et uranifères stratiformes syngenetiques et familières des formations detriques permo-triasiques du Couloir d’Argana (Haut-Atlas occidental, Maroc) . Notes et Mémoires du Service Géologique du Maroc , 33 ( 249 ) : 147 – 177 .
   Google Scholar
• Tixeront , M. 1974 . Carte géologique et minéralogique du Couloir d’Argana, 1/100 000 . Edition du Service Géologique du Maroc, Notes et Mémoires , : 205
   Google Scholar
• Et-Touhami , M. , Olsen , P. E. , Kent , D. V. , Fowell , S. J. and Whiteside , J. H. Tectonostratigraphy, biostratigraphy, and magnetostratigraphy of Late Triassic-Early Jurassic red beds in Morocco . First International Conference & Exhibition, Marrakech, Morocco, Abstracts
   Google Scholar
• Tourani , A. , Lund , J. J. , Benaouiss , N. and Gaupp , R. 2000 . Stratigraphy of Triassic syn-rift deposition in Western Morocco . Zentralblatt für Geologie und Paläontologie Teil I , 9–10 : 1193 – 1215 .
   Google Scholar
• Voigt , S. , Saber , H. , Schneider , J. W. , Hminna , A. , Hmich , D. and Klein , H. 2009 . Large imprints of Hyloidichnus Gilmore, 1927 from the Permian of Morocco in the light of captorrhinid phylogeny and biogeography . : 22 1st International Congress on North African Vertebrate Palaeontology Abstracts
   Google Scholar
• Voigt , S. , Hminna , A. , Saber , H. , Schneider , J. W. and Klein , H. 2010 . Tetrapod footprints from the uppermost level of the Permian Ikakern Formation (Argana basin, western High Atlas, Morocco) . Journal of African Earth Sciences , 57 : 470 – 478 .
   Web of Science ®Google Scholar