Enigmatic Giant Pterosaur Tracks and Associated Ichnofauna from the Cretaceous of Korea: Implication for the Bipedal Locomotion of Pterosaurs

REFERENCES

• Alexander , R. M. 1976 . Estimates of the speeds of dinosaurs . Nature , 261 : 129 – 130 .
   Web of Science ®Google Scholar
• Armstrong , E. A. 1965 . Bird Display and Behavior , 431 New York : Dover Publications Inc. .
   Google Scholar
• Bennett , S. C. 1990 . A pterodactyloid pterosaur pelvis from the Santana Formation of Brazil: Implications for terrestrial locomotion . Journal of Vertebrate Paleontology , 10 : 80 – 85 .
   Google Scholar
• Bennett , S. C. 1997 . Terrestrial locomotion of pterosaurs: A reconstruction based on Pteraichnus trackways . Journal of Vertebrate Paleontology , 17 : 104 – 173 .
   Web of Science ®Google Scholar
• Brunet , M. , Guy , F. Pilbeam , D. 2002 . A new hominid from the Upper Miocene of Chad, Central Africa . Nature , 418 : 145 – 151 .
   PubMed Web of Science ®Google Scholar
• Chang , K. H. 1975 . Cretaceous stratigraphy of southeast Korea . Journal of the Geological Society of Korea , 11 : 1 – 23 .
   Google Scholar
• Chatterjee , S. and Templin , R. J. 2004 . Posture, locomotion, and paleoecology of pterosaurs . Geological Society of America, Special Paper , 376 : 1 – 64 .
   Google Scholar
• Cheong , W. J. 2004 . Report on the survey of dinosaur fossils . Creation Magazine , 138 : 1 – 16 .
   Google Scholar
• Choi , D. K. 1985 . Spores and pollen from the Gyeongsang Supergroup, southeastern Korea and their chronologic and paleoecologic implication . Journal of the Geological Society of Korea , 1 : 33 – 50 .
   Google Scholar
• Choi , H. I. 1986 . Sedimentation and evolution of the Cretaceous Gyeongsang Basin, southeast Korea . Journal of the Geological Society of London , 143 : 29 – 40 .
   Web of Science ®Google Scholar
• Choi , S. J. 1989 . Fossil charophytes from the Nagdong Formation in Seonsangun, Gyeongsangbukdo, Korea . Journal of the Paleontological Society of Korea , 5 : 28 – 38 .
   Google Scholar
• Collini , C. A. 1784 . Sur quleques Zoolithes du Cabinet d’Histoire naturelle de S.A.S.E. Palatine et de Baviére, à Mannheim . Acta Academiae Theodoro-Palatinae Mannheim, Pars Physica , 5 : 58 – 103 .
   Google Scholar
• Cuvier , G. 1801 . Reptile volant . Magasin Encyclopedique, Paris , 9 : 60 – 82 .
   Google Scholar
• Cuvier , G. 1809 . Mémorie sur le squlette fossil d’un reptile volant des environs d’Aichstedt, que quelques naturalistes ont pris pour un oiseau, et don't nous formons un gener de Sauriens, sous le nom de petro-dactyle . Annales du Muséum d’Histoire Naturelle, Paris , 13 : 424 – 437 .
   Google Scholar
• Farlow , J. O. , Pittman , J. G. and Hawthorne , J. M. 1989 . “ Brontopodus birdi, lower Cretaceous footprints from the U. S. Gulf Coastal Plain ” . In Dinosaur Tracks and Traces , Edited by: Gillette , D. D. and Lockley , M. G. 371 – 394 . New York : Cambridge University Press .
   Google Scholar
• Habib , M. B. 2008 . Comparative evidence for quadrupedal launch in pterosaurs . Zitteliana , B28 : 159 – 166 .
   Google Scholar
• Hwang , K. G. , Huh , M. , Lockley , M. G. , Unwin , D. M. and Wright , J. L. 2002 . New pterosaur tracks (Pteraichnidae) from the Late Cretaceous Uhangri Formation, SW Korea . Geological Magazine , 139 : 421 – 435 .
   Web of Science ®Google Scholar
• Kim , B. K. 1969 . A study of several sole marks in the Haman Formation . Journal of the Geological Society of Korea , 5 : 243 – 258 .
   Google Scholar
• Kim , J. Y. , Kim , S. H. , Kim , K. S. and Lockley , M. G. 2006 . The oldest record of webbed bird and pterosaur tracks from South Korea (Cretaceous Haman Formation, Changseon and Sinsu Islands): More evidence of high avian diversity in East Asia . Cretaceous Research , 27 : 56 – 69 .
   Web of Science ®Google Scholar
• Kim , J. Y. , Kim , K. S. , Lockley , M. G. , Yang , S. Y. , Seo , S. J. , Choi , H. I. and Lim , J. D. 2008 . New didactyl dinosaur footprints (Dromaeosauripus hamanensis ichnogen. et ichnosp. nov.) from the Early Cretaceous Haman Formation, south coast of Korea . Palaeogeography, Palaeoclimatology, Palaeoecology , 220 : 72 – 78 .
   Web of Science ®Google Scholar
• Kuban , G. J. 1989 . “ Elongate dinosaur tracks ” . In Dinosaur Tracks and Traces , Edited by: Gillette , D. D. and Lockley , M. G. 57 – 72 . New York : Cambridge University Press .
   Google Scholar
• Lawson , D. A. 1975 . Pterosaur from the latest Cretaceous of West Texas: Discovery of the largest flying creature . Science , 187 : 947 – 948 .
   PubMed Web of Science ®Google Scholar
• Lim , J. D. , Zhou , Z. , Martin , L. D. , Beak , K. S. and Yang , S. Y. 2000 . The oldest known tracks of web-footed birds from the Lower Cretaceous of South Korea . Naturwissenschaften , 87 : 256 – 259 .
   PubMed Web of Science ®Google Scholar
• Lim , S. K. , Yang , S. Y. and Lockey , M. G. 1989 . “ Large dinosaur footprint assemblages from the Cretaceous Jindong Formation of southern Korea ” . Edited by: Gillette , D. D. and Lockley , M. G. 333 – 336 . New York : Cambridge University Press . Dinosaur Tracks and Traces
   Google Scholar
• Lockley , M. G. , Farlow , J. O. and Meyer , C. A. 1994 . Brontopodus and Parabrontopodus ichnogen. Nov. and the significance of wide-and narrow-gauge sauropod trackways . Gaia , 10 : 135 – 145 .
   Google Scholar
• Lockley , M. G. , Harris , J. D. and Mitchell , L. 2008 . A global overview of pterosaur ichnology: Tracksite distribution in space and tome . Zitteliana , B28 : 187 – 200 .
   Google Scholar
• Lockley , M. G. , Houck , K. , Yang , S. Y. , Matsukawa , M. and Lim , S. K. 2006 . Dinosaur-dominated footprint assemblages from the Cretaceous Jindong Formation, Hallyo Haesang National Park area, Goseong Country, South Korea: Evidence and implications . Cretaceous Research , 27 : 70 – 101 .
   Web of Science ®Google Scholar
• Lockley , M. G. , Kim , J. Y. , Kim , K. S. , Kim , S. H. , Matsukawa , M. , Li , R. , Li , J. and Yang , S. Y. 2008 . Minisauripus—the track of a diminutive dinosaur from the Cretaceous of China and South Korea: implication or stratigraphic correlation and theropod foot morphodynamics . Cretaceous Research , 29 : 115 – 130 .
   Web of Science ®Google Scholar
• Lockley , M. G. , Logue , T. J. , Moratalla , J. J. , Hunt , A. P. , Schulz , R. J. and Robinson , J. W. 1995 . The fossil trackway Pteraichnus is pterosaurian, not crocodilian: Implications for the global distribution of pterosaur tracks . Ichnos , 4 : 7 – 20 .
   Google Scholar
• Lockley , M. G. and Wright , J. 2003 . “ Pterosaur swim tracks and other ichnological evidence of behavior and ecology ” . In Evolution and Paleobiology of Pterosaurs , Edited by: Buffetaut , E. Vol. 217 , 297 – 313 . Geological Society of London, Special Publications .
   Google Scholar
• Lockley , M. G. , Yang , S. Y. , Matsukawa , M. , Fleming , F. and Lim , S. K. 1992 . The track record of Mesozoic birds: Evidence and implications . Philosophical Transactions of the Royal Society of Society of London B , 336 : 113 – 134 .
   Web of Science ®Google Scholar
• Mazin , J.-M. , Billon-Briyat , J.-P. , Hantzpergue , P. and LaFaurie , J.-P. 2003 . “ Ichnological evidence for quadrupedal locomotion in pterodactyloid pterosaurs: Trackways from the Late Jurassic of Crayssac (southwestern France) ” . In Evolution and Palaeobiology of Pterosaurs , Edited by: Buffetaut , E. and Mazin , J.-M. Vol. 217 , 283 – 296 . Geological Society of London, Special Publication .
   Google Scholar
• Mickelson , D. L. , Lockley , M. G. , Bishop , J. and Kirkland , J. 2004 . A new pterosaur tracksite from the Jurassic Summerville Formation, near Ferron, Utah . Ichnos , 11 : 125 – 142 .
   Google Scholar
• Padian , K. 1983 . A functional analysis of flying and walking in pterosaurs . Paleobiology , 9 : 218 – 239 .
   Web of Science ®Google Scholar
• Padian , K. 2003 . Pterosaur stance and gait and the interpretation of trackways . Ichnos , 10 : 115 – 126 .
   Google Scholar
• Padian , K. and Olsen , P. E. 1984 . The fossil trackway Pteraichnus: Not pterosaurian, but crocodilian . Journal of Paleontology , 58 : 178 – 184 .
   Web of Science ®Google Scholar
• Pascual Arribas , C. and Sanz Perez , E. 2000 . Huellas de Pterosaurios en el Groupo Oncala (Soria Espana), Pteraichnus palaciei-saenzi, nov. ichnosp . Estudios Geologicos , 56 : 73 – 100 .
   Google Scholar
• Paul , G. S. 1987 . Pterodactyl habits-real and radio-controlled . Nature , 328 : 481
   PubMed Web of Science ®Google Scholar
• Romano , M. , Whyte , M. A. and Jackson , S. J. 2007 . Trackway ratio: A new look at trackway gauge in the analysis of quadrupedal dinosaur trackways and its implications in ichnotaxonomy . Ichnos , 14 : 257 – 270 .
   Google Scholar
• Sarjeant , W. A. S. 1990 . “ A name for the trace of an act: approaches to the nomenclature and classification of vertebrate footprints ” . In Dinosaur Systematics , Edited by: Carpenter , K. and Currie , P. J. 299 – 307 . New York : Cambridge University Press .
   Google Scholar
• Sanchez-Hernadez , B. , Prezewieslik , A. G. and Benton , M. J. 2009 . A reassessment of the Pteraichnus ichnospecies from the early Cretaceous of Spain . Journal of Vertebrate Paleontology , 29 : 487 – 497 .
   Web of Science ®Google Scholar
• Seo , S. J. 1985 . Lower Cretaceous geology and paleontology (Charophyta) of central Kyongsang Basin, Korea. Ph. D. dissertation , 1 – 177 . Korea : Kyungbook National University .
   Google Scholar
• Seo , S. J. 1998 . Dinosaur and pterosaur footprints from the Haman formation, Changsun-myeon, Namhae-gun , 7 Abstract of the 14th Annual Meeting of the Paleontological Society of Korea .
   Google Scholar
• Soemmerring , S. T. von . 1812 . Űber einen Ornithocephalus . 3 : 89 – 158 . Denkschriften der Königlichen Akademie der Wissenschaften zu München
   Google Scholar
• Stokes , W. L. 1957 . Pterodactyl tracks from the Morison Formation . Journal of Paleontology , 31 : 952 – 954 .
   Web of Science ®Google Scholar
• Unwin , D. M. 1987 . Pterosaur locomotion: Joggers or waddlers? . Nature , 327 : 13 – 14 .
   Web of Science ®Google Scholar
• Unwin , D. M. 1989 . “ A predictive method for the identification of vertebrate ichnites and its application to pterosaur track ” . In Dinosaur Tracks and Traces , Edited by: Gillette , D. D. and Lockley , M. G. 259 – 274 . New York : Cambridge University Press .
   Google Scholar
• Unwin , D. M. 1997 . Pterosaur tracks and the terrestrial ability of pterosaurs . Lethaia , 29 : 373 – 386 .
   Web of Science ®Google Scholar
• Wellnhofer , P. 1988 . Terrestrial locomotion in pterosaurs . Historical Biology , 1 : 3 – 16 .
   Google Scholar
• Witton , M. P. and Nash , D. 2008 . A reappraisal of Azhdarchid Pterosaur functional morphology and paleoecology . PLoS One , 3 ( 5 ) : e2271 doi: 10.1371/journal.pone.0002271
   PubMed Web of Science ®Google Scholar
• Wright , J. , Unwin , D. M. , Lockley , M. G. and Rainforth , E. C. 1997 . Pterosaur tracks from the Purbeck Limestone Formation of Dorset, England . Proceedings of the Geologists’ Association , 108 : 39 – 48 .
   Web of Science ®Google Scholar
• Yabe , H. 1905 . Mesozoic plants from Korea . Journal of Science College, Imperial University of Tokyo , 20 : 1 – 76 .
   Google Scholar
• Yang , S. Y. 1976 . On fossils from the Gyeonsang Group-especially on some of the molluscan fauna . Journal of the Geological Society of Korea , 12 : 23 – 30 .
   Google Scholar
• Yang , S. Y. 1982 . On the dinosaur's footprints from the upper Cretaceous Gyeonsang Group, Korea . Journal of the Geological Society of Korea , 18 : 37 – 48 .
   Google Scholar
• Yang , S. Y. , Lockley , M. , Greben , R. and Lim , S. Y. 1995 . Flamingo and duck-like bird tracks from the Late Cretaceous and Early Tertiary: Evidence and implication . Ichnos , 4 : 21 – 34 .
   Google Scholar