First record of a small stegosaur footprint (cf. Stegopodus) from the ?Upper Jurassic-?Lower Cretaceous red beds of the Middle Atlas, Morocco

References

• Bakker, R. T. (1996). The real Jurassic Park: Dinosaurs and habitats at Como Bluff, Wyoming. In: Morales M. (Ed.), The continental Jurassic. The museum of Northern Arizona bulletin (Vol. 60, pp. 35–50).
   Google Scholar
• Charrière, A. (1990). Héritage Hercynien et évolution géodynamique Alpine d‘une chaîne intracontinentale: Le Moyen Atlas au SE de Fès (Maroc) (p. 589). Thèse d‘Etat, Université Paul Sabatier, Toulouse, France.
   Google Scholar
• Charrière, A., & Haddoumi, H. (2016). Les « Couches rouges » continentales jurassico-crétacées des Atlas marocains (Moyen Atlas, Haut Atlas central et oriental): Bilan stratigraphique, paléogéographies successives et cadre géodynamique. Boletín Geológico y Minero, 127 (2/3), 407–430.
   Google Scholar
• Charrière, A., Dépêche, F., Feist, M., Grambast-Fessard, N., Jaffrezo, M., Peybernès, B., & Ramalho, M. (1994). Microfaunes, microflores et paléoenvironnements successifs dans la Formation d’El Mers (Bathonien-? Callovien) du synclinal de Skoura (Moyen-Atlas, Maroc). Geobios, 27 (2), 157–174.
   Google Scholar
• Charrière, A., Ouarhache, D., El Arabi, H. (2011). Le Moyen Atlas (Middle Atlas). In: Michard et al. (Eds.), Notes Et Mémoires du Service Géologique du Maroc (Vol. 559, pp. 11–164). Nouveaux guides géologiques et miniers du Maroc.
   Google Scholar
• Cobos, A., Royo-Torres, R., Luque, L., Alcalá, L., & Mampel, L. (2010). An Iberian stegosaurs paradise: The Villar del Arzobispo Formation (Tithonian-Berriasian) in Teruel (Spain). Palaeogeography, Palaeoclimatology, Paleoecology, 293 (1–2), 223–236.
   Web of Science ®Google Scholar
• du Dresnay, R. (1963). Données stratigraphiques complémentaires sur le Jurassique moyen des synclinaux d’El Mers et de Skoura (Moyen Atlas, Maroc). Bulletin de la Société Géologique de France, 7 (6), 883–900.
   Google Scholar
• Ellenberger, P. (1974). Contribution à la classification des pistes de vertébrés du Trias: Les types du Stormberg d’Afrique du Sud. Palaeovertebrata Mémoire Extaordinaire (pp. 1–142).
   Google Scholar
• Fedan, B. (1993). Évolution géodynamique d’un bassin intraplaque sur décrochements: Le Moyen Atlas (Maroc) durant le Méso-Cénozoique (Vol. 18, p. 144) [Thèse Doctorat d’État 1989]. Trav. Instit. Sci., Rabat, Série Géologie-Géographie-physique.
   Google Scholar
• Frizon de Lamotte, D., Zizi, M., Missenard, Y., Hafid, M., El Azzouzi, M., Maury, R. C., Charrière, A., Taki, Z., Benammi, M., & Michard, A. (2008). The Atlas system. Lecture Notes in Earth Sciences, 116, 33–202.
   Google Scholar
• Gierliński, G. D. (1991). New dinosaur ichnotaxa from the Early Jurassic of the Holy Cross Mountains, Poland. Palaeogeography, Palaeoclimatology, Palaeoecology, 85, 137–148.
   Web of Science ®Google Scholar
• Gierliński, G. D. (1999). Tracks of a large thyreophoran dinosaur from the Early Jurassic of Poland. Acta Palaeontologica Polonica, 44, 231–234.
   Web of Science ®Google Scholar
• Gierliński, G. D., & Pieńkowski, G. (1999). Dinosaur track assemblages from the Hettangian of Poland. Geological Quarterly, 43, 329–346.
   Google Scholar
• Gierliński, G. D., & Sabath, K. (2002). A probable stegosaurian track from the Late Jurassic of Poland. Acta Palaeontologica Polonica, 47(3), 561–564.
   Google Scholar
• Gierliński, G. D., & Sabath, K. (2008). Stegosaurian footprints from the Morrison Formation of Utah and their implication for interpreting other ornitischian tracks. Oryctos, 8, 29–46.
   Google Scholar
• Gierliński, G. D., Szrek, P., & Lockley, M. G. (2010). New finds of dinosaur tracks in the Morrison Formation of Moab area, Utah, USA. Earth Science Frontiers, 17, 167–168.
   Google Scholar
• Gierliński, G. D., Menducki, P., Janiszewska, K., Wicik, I., & Bosczarowski, A. (2009). A preliminary report on dinosaur track assemblages from the Middle Jurassic of the Imilchil area, Morocco. Geological Quarterly, 53(4), 477–482.
   Web of Science ®Google Scholar
• Hadri, M., & Pérez-Lorente, F. (2012). Historia de yacimientos con huellas de dinosaurio, desde su descubrimiento hasta su primer estudio. Alrededores de El Mers (Marruecos). Zubía, 30, 93–140.
   Google Scholar
• Han, F., Zheng, W., Hu, D., Xu, X., & Barrett, P. M. (2014). A new basal ankylosaurid (Dinosauria: Ornithischia) from the Lower Cretaceous Jiufotang Formation of Liaoning Province, China. PLoS One, 9(8), e104551.
   PubMed Web of Science ®Google Scholar
• Klein, H., Gierliński, G. D., Oukassou, M., Saber, H., Lallensack, J. N., Lagnaoui, A., Hmmina, A., & Charrière, A. (2022). Theropod and ornithischian dinosaur track assemblages from Middle to ?Late Jurassic deposits of the Central High Atlas, Morocco. Historical Biology, 35, 320–346.
   Web of Science ®Google Scholar
• Kuhn, O. (1958). Die Fährten der vorzeitlichen Amphibien und Reptilien (p. 64). Verlagshaus Meisenbach.
   Google Scholar
• Lapparent, A. F. de. (1955). Etude paléontologique des vertébrés du Jurassique d’El Mers (Moyen Atlas). Notes et Mémoires du Service Géologique du Maroc, 124, 1–36.
   Google Scholar
• Li, J., Lockley, M. G., Zhang, Y. G., Hu, S. M., Matsukawa, M., & Bai, Z. Q. (2012). An important omithischian tracksite in the Early Jurassic of the Shenmu Region, Shaanxi, China. Acta Geologica Sinica - English Edition, 86, 1–10. https://doi.org/10.1111/j.1755-6724.2012.00606.x
   Web of Science ®Google Scholar
• Lockley, M. G., & Hunt, A. P. (1998). A probable stegosaur track from the Morrison Formation of Utah. Modern Geology, 23, 331–342.
   Google Scholar
• Lockley, M. G., McCrea, R. T., & Buckley, L. G. (2015). A review of dinosaur track occurrences from the Morrison Formation in the type area around Dinosaur Ridge. Palaeogeography, Palaeoclimatology, Paleoecology, 433, 10–19.
   Web of Science ®Google Scholar
• Lockley, M. G., Gierliński, G. D., Matthews, N., Xing, L., Foster, J., & Cart, K. (2017). New dinosaur track occurrences from the Upper Jurassic Salt Wash Member (Morrison Formation) of southeastern Utah: Implications for thyreophoran trackmaker distribution and diversity. Palaeogeography, Palaeoclimatology, Paleoecology, 470, 116–121.
   Web of Science ®Google Scholar
• Maidment, S. C., Raven, T. J., Ouarhache, D., & Barrett, P. M. (2020). North Africa’s first stegosaur: Implications for Gondwanan thyreophoran dinosaur diversity. Gondwana Research, 77, 82–97.
   Web of Science ®Google Scholar
• Maidment, S. C., Strachan, S. J., Ouarhache, D., Scheyer, T. M., Brown, E. E., Fernandez, V., Johanson, Z., Raven, T. J., & Barrett, P. M. (2021). Bizarre dermal armour suggests the first African ankylosaur. Nature Ecology and Evolution, 5, 1576–1581. https://doi.org/10.1038/s41559-021-01553-6
   PubMed Web of Science ®Google Scholar
• Marchetti, L., Belvedere, M., Voigt, S., Klein, H., Castanera, D., Díaz-Martínez, I., Marty, D., Xing, L., Feola, S., Melchor, R. N., & Farlow, J. O. (2019). Defining the morphological quality of fossil footprints. Earth-Science Reviews, 193, 109–145. https://doi.org/10.1016/j.earscirev.2019.04.008
   Web of Science ®Google Scholar
• Meyer, C. A., & Thüring, B. (2004). The first dinosaur footprints from the Middle Jurassic (Bathonian-Callovian) of the Middle Atlas mountains (Morocco) (p. 20). 2nd Meeting of European Vertebrate Palaeontologist. Brno, Abstracts with Programm.
   Google Scholar
• Meyer, C. A., & Thüring, B. (2005, November 18-19). Mind the “Middle Jurassic” gap. Bone versus track record in dinosaurs (pp. 59–60). 3rd Swiss Geoscience Meeting, Zurich.
   Google Scholar
• Milàn, J. (2011). New theropod, thyreophoran, and small sauropod tracks from the Middle Jurassic Bagå Formation, Bornholm, Denmark. Bulletin of the Geological Society of Denmark, 59, 51–59. https://doi.org/10.37570/bgsd-2011-59-06
   Web of Science ®Google Scholar
• Mossbrucker, M., Bakker, R., Lockley, M. G., Gierliński, G. D., & Hups, K. (2008). Probable stegosaur tracks from the Jurassic Morrison Formation, Colorado and Utah. Dinosaur Ridge Annual Report, 20(5), 26–29.
   Google Scholar
• Ouarhache, D. (2002). Sédimentation et volcanismes (effusif et explosif) associés au rifting triasique et infraliasique dans le Moyen Atlas sud-occidental et la Haute Moulouya (Maroc) (p. 282). Thèse Sci., Univ. Mohammed V, Rabat, deposed in S.G.F, Paris.
   Google Scholar
• Ouarhache, D., Charrière, A., Chalot-Prat, F., & El Wartiti, M. (2012). Chronologie et modalités du rifting triasico-liasique à la marge sud-ouest de la Téthys alpine (Moyen Atlas et Haute Moulouya, Maroc); corrélations avec le rifting atlantique: Simultanéité et diachronisme. Bulletin de la Société Géologique de France, 183 (3), 233–249.
   Google Scholar
• Oukassou, M., Charrière, A., Lagnaoui, A., Gibb, S., Michard, A., & Saddiqi, O. (2016). First occurrence of the ichnogenus Selenichnites from the Middle Jurassic strata of the Skoura syncline (Middle Atlas, Morocco); palaeoeological and palaeoenvironmental context. Comptes Rendus Palevol, 15 (5), 461–471. https://doi.org/10.1016/j.crpv.2015.09.013
   Web of Science ®Google Scholar
• Oukassou, M., Boumir, K., Benshili, K., Ouarhache, D., Lagnaoui, A., & Charrière, A. (2019). The Tichoukt massif: A geotouristic play in the folded Middle Atlas (Morocco). Geoheritage, 11 (2), 371–379. https://doi.org/10.1007/s12371-018-0287-y
   Web of Science ®Google Scholar
• Piñuela, L., Garcia-Ramos, J. C., & Lires, J. (2002). Comportamiento gregario de ornitópodos en el Jurásico de Asturias. In F. Perez-Lorente (Ed.), Dinosaurios y otros reptiles mesozoicos en España. Resúmenes de las comunicaciones ponencias y panels, Logroño (Vol. 27, pp. 44–45).
   Google Scholar
• Pratt, J. R., Barbeau, D. L., Jr., Garver, J. I., Emran, A., & Izykowski, T. M. (2015). Detrital zircon geochronology of Mesozoic sediments in the Rif and Middle Atlas belts of Morocco: Provenance constraints and refinement of the West African signature. The Journal of Geology, 123, 177–200. https://doi.org/10.1086/681218
   Web of Science ®Google Scholar
• Sternberg, C. M. (1932). Dinosaur tracks from Peace River, British Columbia. Bulletin of the National Museum of Canada, 68, 59–85.
   Google Scholar
• Termier, H., Gubler, J., & de Lapparent, A. F. (1940). Le Bathonien de la région d‘El Mers: Reptiles et poisons due Bathonien D'El mers. Compte rendus hebdomadaires des séances de l‘académie des sciences (pp. 1–2).
   Google Scholar
• Thulborn, R. A., (1990). Dinosaur tracks (p. 410). Chapman & Hall.
   Google Scholar
• Whyte, M., & Romano, M. (1994). Probable sauropod footprints from the Middle Jurassic of Yorkshire, England. Gaia, 10, 15–26.
   Google Scholar
• Xing, L., Lockley, M. G., McCrea, R. T., Gierliński, G. D., Buckley, L. G., Zhang, J., Qi, L., & Jia, C. (2013). First record of Deltapodus tracks from the Early Cretaceous of China. Cretaceous Research, 42, 55–65. http://dx.doi.org/10.1016/j.cretres.2013.01.006
   Web of Science ®Google Scholar
• Xing, L., Lockley, M. G., Klein, H., Gierliński, G. D., Ye, Y., Zhang, J., Persons, W. S. IV., & Wang, T. (2019). First thyreophoran type tracks from the Middle Jurassic Chuangjie Formation of Yunnan Province, China. Ichnos, 26, 8–15. https://doi.org/10.1080/10420940.2017.1366904
   Web of Science ®Google Scholar
• Xing, L., Lockley, M. G., Scott Persons, W., Klein, H., Romilio, A., Wang, D., & Wang, M. (2021). Stegosaur track assemblage from Xinjiang, China, featuring the smallest known stegosaur record. Palaios, 36(2), 68–76. https://doi.org/10.2110/palo.2020.036
   Web of Science ®Google Scholar
• Zafaty, O., Oukassou, M., Pereda-Suberbiola, X., Riguetti, F., Company, J., Bendrioua, S., Tabuce, R., & Charrière, A. (2022, May 20–22). New remains of Thyreophoran dinosaurs (Ornithischia) from the Middle Atlas Jurassic red beds of Morocco. 4èmes Journées Jeunes Chercheurs en Géosciences (JJCG2022), Tétouan, Maroc.
   Google Scholar