Similar tree seedling responses to shrubs and to simulated environmental changes at Pyrenean and subarctic treelines

References

• Akhalkatsi , M , Abdaladze , O , Nakhutsrishvili , G and Smith , W. 2006 . Facilitation of seedling microsites by Rhododendron caucasicum extends the Betula litwinowii Alpine treeline, Caucasus Mountains, Republic of Georgia . Arctic, Antarctic, and Alpine Research , 38 : 481 – 488 .
   Web of Science ®Google Scholar
• Alados , C , Komac , B , Bueno , CG , Gartzia , M , Escós , J , Gómez García , D , García-González , R , Fillat , F , Camarero , JJ and Herrero , J . 2011 . “ Modelización de la matorralización de los pastos del Parque Nacional de Ordesa y Monte Perdido y su relación con el cambio global ” . In Proyectos de investigación en parques nacionales 2007–2010 , Edited by: Ramírez , L and Asensio , B . Madrid (Spain): Organismo Autónomo Parques Nacionales .
   Google Scholar
• Alftine , K and Malanson , G. 2004 . Directional positive feedback and pattern at an alpine tree line . Journal of Vegetation Science , 15 : 3 – 12 .
   Web of Science ®Google Scholar
• Ameztegui , A and Coll , A. 2011 . Tree dynamics and co-existence in the montane sub-alpine ecotone: the role of different light-induced strategies . Journal of Vegetation Science , 22 : 1049 – 1061 .
   Web of Science ®Google Scholar
• Ameztegui , A , Brotons , LL and Coll , LL. 2010 . Land-use changes as major drivers of mountain pine (Pinus uncinata Ram.) . expansion in the Pyrenees. Global Ecology and Biogeography , 19 : 632 – 641 .
   Web of Science ®Google Scholar
• Andrews , C , Dick , J , Jonasson , C and Callaghan , TV. 2011 . Assessment of biological and environmental phenology at a landscape level from 30 years of fixed date repeat photography in Northern Sweden . Ambio , 40 : 600 – 609 .
   PubMed Web of Science ®Google Scholar
• Anschlag , K , Broll , G and Holtmeier , FK. 2008 . Mountain birch seedlings in the treeline ecotone, subarctic Finland: variation in above- and below-ground growth depending on microtopography . Arctic, Antarctic, and Alpine Research , 40 : 609 – 616 .
   Web of Science ®Google Scholar
• Batllori , E and Gutiérrez , E. 2008 . Regional treeline dynamics in response to global change in the Pyrenees . Journal of Ecology , 96 : 1275 – 1288 .
   Web of Science ®Google Scholar
• Batllori , E , Camarero , JJ and Gutiérrez , E. 2010 . Current regeneration patterns at the treeline in the Pyrenees indicate similar recruitment processes irrespective of the past disturbance regime . Journal of Biogeography , 37 : 1938 – 1950 .
   Web of Science ®Google Scholar
• Batllori , E , Camarero , J , Ninot , JM and Gutiérrez , E. 2009 . Seedling recruitment, survival and facilitation in alpine Pinus uncinata tree line ecotones . Implications and potential responses to climate warming. Global Ecology and Biogeography Letters , 18 : 460 – 472 .
   Web of Science ®Google Scholar
• Bokhorst , S , Bjerke , J , Davey , M , Taulavuori , K , Taulavuori , E and Laine , K. 2010 . Impacts of extreme winter warming events on plant physiology in a sub-Arctic heath community . Physiologia Plantarum , 140 : 128 – 140 .
   PubMed Web of Science ®Google Scholar
• Bokhorst , S , Bjerke , J , Tømmervik , H , Callaghan , TV and Phoenix , GK. 2009 . Winter warming events damage sub-Arctic vegetation: consistent evidence from an experimental manipulation and a natural event . Journal of Ecology , 97 : 1408 – 1415 .
   Web of Science ®Google Scholar
• Bokhorst , S , Bjerke , JW , Bowles , FW , Melillo , J , Callaghan , TV and Phoenix , GK. 2008 . Impacts of extreme winter warming in the sub-Arctic: growing season responses of dwarf shrub heathland . Global Change Biology , 14 : 2603 – 2612 .
   Web of Science ®Google Scholar
• Brooker , R and Callaghan , TV. 1998 . The balance between positive and negative plant interactions and its relationship to environmental gradients: a model . Oikos , 81 : 196 – 206 .
   Web of Science ®Google Scholar
• Cairns , D and Moen , J. 2004 . Herbivory influences tree lines . Journal of Ecology , 92 : 1019 – 1024 .
   Web of Science ®Google Scholar
• Callaghan , TV , Bergholm , F , Christensen , T , Jonasson , C , Kokfelt , U and Johansson , M. 2010 . A new climate era in the sub-Arctic: accelerating climate changes and multiple impacts . Geophysical Research Letters , 37 : L14705
   Web of Science ®Google Scholar
• Cantegrel , R. 1983 . Le pin à crochets pyrénéen: biologie, biochimie, sylviculture . Acta Biologica Montana , 2 : 87 – 330 .
   Google Scholar
• Canut , J. 2001 . “ Mountain gallinaceous (partridge, ptarmigan and capercaillie) and forest management ” . In Conservación de la biodiversidad, fauna vertebrada y gestión forestal , 2nd , Edited by: Camprodon , J and Plana , E . Barcelona (Spain): Centre Tecnològic Forestal de Catalunya and Universitat de Barcelona .
   Google Scholar
• Canut , J. 1991 . El gall fer a Catalunya: situació, actuacions i conservació . Medi ambient , 2 : 74 – 78 .
   Google Scholar
• Carreras , J , Carrillo , E , Masalles , R , Ninot , J , Soriano , I and Vigo , J. 1996 . Delimitation of the supra-forest zone in the Catalan Pyrenees . Bulletin de la Societé linnéenne de Provence , 47 : 27 – 36 .
   Google Scholar
• Chauchard , S , Beilhe , F , Denis , N and Carcaillet , C. 2010 . An increase in the upper tree-limit of silver fir (Abies alba Mill.) in the Alps since the mid-20th century: a land-use change phenomenon . Forest Ecology and Management , 259 : 1406 – 1415 .
   Web of Science ®Google Scholar
• Cornelissen , JHC , Werger , MJA , Castro-Díez , P , van Rheenen , JWA and Rowland , AP. 1997 . Foliar nutrients in relation to growth, allocation and leaf traits in seedlings of a wide range of woody plant species and types . Oecologia , 111 : 460 – 469 .
   PubMed Web of Science ®Google Scholar
• Cunill , R. 2010 . “ Estudi interdisciplinari de l'evolució del límit superior del bosc durant el període holocènic a la zona de Plaus de Boldís-Montarenyo ” . In Pirineu central català. Pedoantracologia, palinologia, carbons sedimentaris i fonts documentals [Ph.D. thesis] , Barcelona ( , , Catalonia)] : Universitat Autònoma de Barcelona .
   Google Scholar
• Dalen , L and Hofgaard , A. 2005 . Differential regional treeline dynamics in the Scandes Mountains . Arctic, Antarctic, and Alpine Research , 37 : 284 – 296 .
   Web of Science ®Google Scholar
• Daniels , L and Veblen , T. 2004 . Spatiotemporal influences of climate on altitudinal treeline in northern Patagonia . Ecology , 85 : 1284 – 1296 .
   Web of Science ®Google Scholar
• Díaz-de-Quijano , M , Peñuelas , J and Ribas , À. 2009 . Increasing interannual and altitudinal ozone mixing ratios in the Catalan Pyrenees . Atmospheric Environment , 43 : 6049 – 6057 .
   Web of Science ®Google Scholar
• Elias , SA. 2001 . “ Paleoecology and late Quaternary environments of the Colorado Rockies ” . In Structure and Function of an Alpine Ecosystem, Niwot Ridge, Colorado , Edited by: Bowman , WD and Seastedt , TR . 285 – 303 . Oxford ( , , UK) : Oxford University Press .
   Google Scholar
• Elliott , G. 2012 . The role of thresholds and fine-scale interactions in driving upper treeline dynamics in the Bighorn Mountains, Wyoming . Physical Geography , 33 : 129 – 145 .
   Web of Science ®Google Scholar
• Elliott , G. 2011 . Influences of 20th-century warming at the upper tree line contingent on local-scale interactions: evidence from a latitudinal gradient in the Rocky Mountains, USA . Global Ecology and Biogeography , 20 : 46 – 57 .
   Web of Science ®Google Scholar
• Elliott , G and Kipfmueller , K. 2010 . Multi-scale influences of slope aspect and spatial pattern on ecotonal dynamics at upper treeline in the southern Rocky Mountains, USA . Arctic, Antarctic, and Alpine Research , 42 : 45 – 56 .
   Web of Science ®Google Scholar
• Elmendorf , SC , Henry , GHR , Hollister , RD , Björk , RG , Boulanger-Lapointe , N , Cooper , EJ and Cornelissen , JHC . 2012 . Plot-scale evidence of tundra vegetation change and links to recent summer warming . Nature Climate Change , l2 : 453 – 457 .
   Web of Science ®Google Scholar
• Esteban , P , Prohom , MJ , Aguilar , E and Mestre , O. 2009 . Evolució recent de la temperatura i de la precipitació a Andorra (1934–2008): resultats anuals i estacionals , Andorra : Centre d'Estudis de la Neu i de la Muntanya d'Andorra de l'Institut d'Estudis Andorrans (CENMA) .
   Google Scholar
• Frich , P , Alexander , LV , Della-Marta , P , Gleason , B , Haylock , M , Klein Tank , AMG and Peterson , T. 2002 . Observed coherent changes in climatic extremes during the second half of the twentieth century . Climate Research , 19 : 193 – 212 .
   Web of Science ®Google Scholar
• Gehrig-Fasel , J , Guisan , A and Zimmermann , N. 2007 . Tree line shifts in the Swiss Alps . climate change or land abandonment? Journal of Vegetation Science , 18 : 571 – 582 .
   Google Scholar
• Germino MJ, Smith WK. 1999. Sky exposure, crown architecture, and low-temperature photoinhibition in conifer seedlings at alpine treeline. Plant Cell and Environment 22:407–415.
   Google Scholar
• Germino , MJ , Smith , WK and Resor , AC. 2002 . Conifer seedling distribution and survival in an alpine-treeline ecotone . Plant Ecology , 162 : 157 – 168 .
   Web of Science ®Google Scholar
• Grace , J. 1990 . Cuticular water loss unlikely to explain tree-line in Scotland . Oecologia , 84 : 64 – 68 .
   PubMed Web of Science ®Google Scholar
• Grace , J , Berninger , F and Nagy , L. 2002 . Impacts of climate change on the tree line . Annals of Botany , 90 : 537 – 544 .
   PubMed Web of Science ®Google Scholar
• Grau , O , Ninot , JM , Blanco-Moreno , JM , van Logtestijn , RSP , Cornelissen , JHC and Callaghan , TV. 2012 . Shrub-tree interactions and environmental changes drive treeline dynamics in the Subarctic . Oikos , 121 : 1680 – 1690 .
   Web of Science ®Google Scholar
• Grime , JP. 2001 . Plant strategies, vegetation processes and ecosystem properties , Chichester (UK) : John Wiley and Sons .
   Google Scholar
• Hadley , JL and Smith , WK. 1989 . Wind erosion of leaf surface wax in alpine timberline conifers . Arctic and Alpine Research , 21 : 392 – 398 .
   Web of Science ®Google Scholar
• Hallinger , M , Manthey , M and Wilmking , M. 2010 . Establishing a missing link: warm summers and winter snow cover promote shrub expansion into alpine tundra in Scandinavia . New Phytologist , 18 : 890 – 899 .
   Web of Science ®Google Scholar
• Harsch , MA , Hulme , PE , McGlone , MS and Duncan , RP. 2009 . Are treelines advancing? A global meta-analysis of treeline response to climate change . Ecology Letters , 12 : 1040 – 1049 .
   PubMed Web of Science ®Google Scholar
• Hedenås , H , Olsson , H , Jonasson , C , Bergstedt , J , Dahlberg , U and Callaghan , TV. September 2011 . “ Tree growth, biomass and vegetation changes over a thirteen-year period in the Swedish sub-Arctic ” . In Multi-decadal changes in tundra environments and ecosystems: the International Polar Year Back to the Future project , Edited by: Callaghan , TV and Tweedie , CE . Vol. 40 , September , 672 – 682 . Ambio .
   Google Scholar
• Henríquez , J and Lusk , C. 2005 . Facilitation of Nothofagus antarctica (Fagaceae) seedlings by the prostrate shrub Empetrum rubrum (Empetraceae) on glacial moraines in Patagonia . Austral Ecology , 30 : 877 – 882 .
   Web of Science ®Google Scholar
• Holtmeier , F and Broll , G. 2005 . Sensitivity and response of northern hemisphere altitudinal and polar treelines to environmental change at landscape and local scales . Global Ecology and Biogeography Letters , 14 : 395 – 410 .
   Web of Science ®Google Scholar
• Holtmeier , FK , Broll , G , Muterthies , A and Anschlag , K. 2003 . Regeneration of trees in the treeline ecotone: northern Finnish Lapland . Fennia , 181 : 103 – 128. .
   Google Scholar
• [IPCC] Intergovernmental Panel on Climate Change . 2007 . “ Climate change 2007: the physical science basis ” . In Contribution of working group I to the fourth assessment report of the Intergovernmental Panel on Climate Change , Edited by: Solomon , S , Qin , D , Manning , M , Chen , Z , Marquis , M , Averyt , KB , Tignor , M and Miller , HL . Cambridge ( , , UK) : Cambridge University Press .
   Google Scholar
• Jepsen , J , Kapari , L , Hagen , SB , Vindstad , OPL , Nilssen , AC and Ims , RA. 2011 . Rapid northwards expansion of a forest insect pest attributed to spring phenology matching with sub-Arctic birch . Global Change Biology , 17 : 2071 – 2083 .
   Web of Science ®Google Scholar
• Karlsson , P and Nordell , K. 1996 . Effects of soil temperature on the nitrogen economy and growth of mountain birch seedlings near its presumed low temperature distribution limit . Écoscience , 3 : 183 – 189 .
   Web of Science ®Google Scholar
• Komac , B , Kefi , S , Nuche , P , Escós , J and Alados , CL. 2013 . Modeling shrub encroachment in subalpine grasslands under different environmental and management scenarios . Journal of Environmental Management , 121 : 160 – 169 .
   PubMed Web of Science ®Google Scholar
• Körner , C. 2012 . Alpine treelines. Functional ecology of the global high elevation tree limits , Basel , , Switzerland : Springer .
   Google Scholar
• Körner , C and Paulsen , J. 2004 . A world-wide study of high elevation treeline temperatures . Journal of Biogeography , 31 : 713 – 732 .
   Web of Science ®Google Scholar
• Kullman , L. 2002 . Rapid recent range-margin rise of tree and shrub species in the Swedish Scandes . Journal of Ecology , 90 : 68 – 77 .
   Web of Science ®Google Scholar
• Kullman , L and Öberg , L. 2009 . Post-Little Ice Age tree line rise and climate warming in the Swedish Scandes: a landscape ecological perspective . Journal of Ecology , 97 : 415 – 429 .
   Web of Science ®Google Scholar
• Lasanta , T. 1990 . “ Diversidad de usos e integración especial en la gestión tradicional del territorio en las montañas de Europa Occidental ” . In Geoecología de las áreas de montaña , Edited by: García , JM . Logroño , , Spain : Geoforma Ediciones .
   Google Scholar
• Llambí , LD , Puentes , J and García-Nuñez , C. 2013 . “ Spatial relationships and population structure of a dominant tree along a treeline ecotone in the tropical Andes: interactions at landscape and plant-neighbourhood scales ” . In Plant Ecology & Diversity 6 343 – 353 .
   Google Scholar
• Maestre , F , Callaway , R , Valladares , F and Lortie , C. 2009 . Refining the stress-gradient hypothesis for competition and facilitation in plant communities . Journal of Ecology , 97 : 199 – 205 .
   Web of Science ®Google Scholar
• Marion , GM , Henry , GHR , Freckman , DW , Johnstone , J , Jones , G , Jones , MH , Lévesque , E , Molau , U , MØlgaard , P and Parsons , AN . 1997 . Open-top designs for manipulating field temperature in high-latitude ecosystems . Global Change Biology , 3 : 20 – 32 .
   Web of Science ®Google Scholar
• Motta , R and Nola , P. 2001 . Growth trends and dynamics in sub-alpine forest stands in the Varaita Valley (Piedmont, Italy) and their relationships with human activities and global change . Journal of Vegetation Science , 12 : 219 – 230 .
   Web of Science ®Google Scholar
• Myers-Smith , I , Forbes , B , Wilmking , M , Hallinger , M , Lantz , T , Blok , D , Tape , K , Macias-Fauria , M , Sass-Klaassen , U Lévesque , E . 2011 . Shrub expansion in tundra ecosystems: dynamics, impacts and research priorities . Environmental Research Letters , 6 : 045509
   Web of Science ®Google Scholar
• Nagy , J , Nagy , L , Legg , CJ and Grace , J. 2013 . The stability of the Pinus sylvestris treeline in the Cairngorms, Scotland over the last millennium . Plant Ecology & Diversity , 6 : 7 – 19 .
   Web of Science ®Google Scholar
• Naito , A and Cairns , D. 2011 . Patterns and processes of global shrub expansion . Progress in Physical Geography , 35 : 423 – 442. .
   Web of Science ®Google Scholar
• Neuner , G. 2007 . “ Frost resistance at the upper timberline ” . In Trees at their upper limit. Tree life limitation at the alpine timberline , Edited by: Wieser , G and Tausz , M . 171 – 180 . Dordrecht , , The Netherlands : Springer .
   Google Scholar
• Ninyerola M, Pons X, Roure JM. 2000. A methodological approach of climatological modelling of air temperature and precipitation through GIS techniques. International Journal of Climatology 20:1823–1841. The Digital climatic atlas of Catalonia is available online at (accessed 2011) http://www.opengis.uab.cat/acdc/en_index.htm (http://www.opengis.uab.cat/acdc/en_index.htm)
   Google Scholar
• Ovaska , J , Nilsen , J , Wielgolaski , FE , Kauhanen , H , Partanen , R , Neuvonen , S , Kapari , L , Skre , O and Laine , K. 2005 . “ Phenology and performance of mountain birch provenances in transplant gardens: latitudinal, altitudinal and oceanity-continentality gradients ” . In Plant ecology, herbivory, and human impact in Nordic mountain birch forests , Edited by: Wielgolaski , FE . 99 – 115 . Berlin , , Germany : Springer .
   Google Scholar
• Pornon , A , Escaravage , N , Till-Bottraud , I and Doche , B. 1997 . Variation of reproductive traits in Rhododendron ferrugineum L. (Ericaceae) populations along a successional gradient . Plant Ecology , 130 : 1 – 11 .
   Web of Science ®Google Scholar
• R Development Core Team . 2011 . R: a language and environment for statistical computing , Vienna , , Austria : R Foundation for Statistical Computing .
   Google Scholar
• Resler , L , Butler , D and Malanson , G. 2005 . Topographic shelter and conifer establishment and mortality in an alpine environment, Glacier National Park, Montana . Physical Geography , 26 : 112 – 125 .
   Web of Science ®Google Scholar
• Rundqvist , S , Hedenås , H , Sandström , A , Emanuelsson , U , Eriksson , H , Jonasson , C and Callaghan , TV. September 2011 . “ Tree and shrub layer expansion over the past 34 years at the tree-line in Abisko, sub-Arctic ” . In Multi-decadal changes in tundra environments and ecosystems: the International Polar Year Back to the Future project , Edited by: Callaghan , TV and Tweedie , CE . Vol. 40 , September , 683 – 692 . Ambio .
   Google Scholar
• Rupp , TS , Chapin , FS I II and Starfield , AM. 2001 . Modelling the influence of topographic barriers on treeline at the forest tundra ecotone in Northwestern Alaska . Climate Change , 48 : 399 – 416 .
   Web of Science ®Google Scholar
• Rustad , LE , Campbell , JL , Marion , GM , Norby , RJ , Mitchell , MJ , Hartley , AE , Cornelissen , JHC and Gurevitch , J. 2001 . A meta-analysis of the response of soil respiration, net nitrogen mineralization, and aboveground plant growth to experimental ecosystem warming . Oecologia , 126 : 543 – 562 .
   PubMed Web of Science ®Google Scholar
• Sigrun , A , Hofgaard , A and Soderstrom , L. 2011 . Contrasting climate- and land-use-driven tree encroachment patterns of subarctic tundra in northern Norway and the Kola Peninsula . Canadian Journal of Forest Research. , 41 : 437 – 449 .
   Web of Science ®Google Scholar
• [SMC] Servei Meteorològic de Catalunya. 2012. Available online at (accessed 2012) http://www.meteo.cat (http://www.meteo.cat)
   Google Scholar
• Smith , W , Germino , M , Hancock , T and Johnson , D. 2003 . Another perspective on altitudinal limits of alpine timberlines . Tree Physiology , 23 : 1101 – 1112 .
   PubMed Web of Science ®Google Scholar
• Sullivan , P and Sveinbjörnsson , B. 2010 . Microtopographic control of treeline advance in Noatak National Preserve, Northwest Alaska . Ecosystems , 13 : 275 – 285 .
   Web of Science ®Google Scholar
• Stueve , K , Cerney , D , Rochefort , R and Kurth , L. 2009 . Post-fire tree establishment patterns at the alpine treeline ecotone: Mount Rainier National Park, Washington, USA . Journal of Vegetation Science , 20 : 107 – 120 .
   Web of Science ®Google Scholar
• Stueve , K , Isaacs , R , Tyrrell , L and Densmore , R. 2011 . Spatial variability of biotic and abiotic tree establishment constraints across a treeline ecotone in the Alaska Range . Ecology , 92 : 496 – 506 .
   PubMed Web of Science ®Google Scholar
• Sturm , M , Racine , C and Tape , K. 2001 . Increasing shrub abundance in the Arctic . Nature , 411 : 546 – 7 .
   PubMed Web of Science ®Google Scholar
• Sveinbjörnsson , B. 2001 . “ Performance of mountain birch in relation to its treeline in the Torneträsk area, northern Sweden ” . In Nordic mountain birch ecosystems , Edited by: Wielgolaski , FE . 173 – 183 . Paris , , France : UNESCO .
   Google Scholar
• Tape , K , Sturm , M and Racine , C. 2006 . The evidence for shrub expansion in Northern Alaska and the Pan-Arctic . Global Change Biology , 12 : 686 – 702 .
   Web of Science ®Google Scholar
• Tenow , O. 1996 . Hazards to a mountain birch forest: Abisko in perspective . Ecological Bulletins , 45 : 104 – 114 .
   Google Scholar
• Tinner , W and Theurillat , J. 2003 . Uppermost limit, extent, and fluctuations of the timberline and treeline ecocline in the Swiss Central Alps during the past 11,500 years . Arctic, Antarctic, and Alpine Resesarch , 35 : 158 – 169 .
   Web of Science ®Google Scholar
• Tomback , DF and Resler , LM. 2007 . Invasive pathogens at alpine treeline: consequences for treeline dynamics . Physical Geography , 28 : 397 – 418 .
   Web of Science ®Google Scholar
• Tømmervik , H , Johansen , B , Tombre , I , Thannheiser , D , Hogda , K and Gaare , E. 2004 . Vegetation changes in the Nordic mountain birch forest: The influence of grazing and climate change . Arctic, Antarctic, and Alpine Research , 36 : 323 – 332 .
   Web of Science ®Google Scholar
• Van Bogaert , R , Jonasson , C , De Dapper , M and Callaghan , TV. 2009 . Competitive interaction between aspen and birch moderated by invertebrate and vertebrate herbivores and climate warming . Plant Ecology & Diversity , 2 : 221 – 232 .
   Web of Science ®Google Scholar
• Van Bogaert , R , Haneca , K , Hoogesteger , J , Jonasson , C , De Dapper , M and Callaghan , TV. 2011 . A century of tree line changes in sub-Arctic Sweden show local and regional variability and only a minor role of 20th century climate warming . Journal of Biogeography , 38 : 907 – 921 .
   Web of Science ®Google Scholar
• Wheeler , J , Hermanutz , L and Marino , P. 2011 . Feathermoss seedbeds facilitate black spruce seedling recruitment in the forest-tundra ecotone (Labrador, Canada) . Oikos , 120 : 1263 – 1271 .
   Web of Science ®Google Scholar
• Wielgolaski , F. , Kankaanpää , S and Muller-Wille , L . 2002 . “ Northern timberlines forests: Environmental and socio-economic issues and concerns ” . In The Finnish Forest Research Institute , Edited by: Susiluoto , P and Sutinen , ML . Vol. 862 , 128 – 132 . 266 – 271 . Kolari : Research Papers .
   Google Scholar
• Wilmking , M and Juday , G. 2005 . Longitudinal variation of radial growth at Alaska's northern treeline-recent changes and possible scenarios for the 21st century . Global and Planetary Change , 47 : 282 – 300 .
   Web of Science ®Google Scholar
• Wolf , A , Larsson , K and Callaghan , TV. 2008 . Future vegetation changes in the Barents Region . Climatic Change , 87 : 51 – 73 .
   Web of Science ®Google Scholar
• Zhang Y, Wilmking M. 2010. Divergent growth responses and increasing temperature limitation of Qinghai spruce growth along an elevation gradient at the northeast Tibet Plateau. Forest Ecology and Management 260:1076–1082.
   Google Scholar