The padloper’s tortuous path (Chelonia: Testudinidae): Two genera, not one

References

• Bates, M.F., W.R. Branch, A.M. Bauer, M. Burger, J. Marais, G.J. Alexander & M.S. de Villiers (Eds). 2014. Atlas and Red List of the Reptiles of South Africa, Lesotho and Swaziland. Suricata 1. South African National Biodiversity Institute, Pretoria.
   Google Scholar
• Bour, R. 1988. Taxonomic and nomenclatural status of Homopus signatus (Gmelin, 1789): Reptilia-Chelonii. J. Herpetol. Ass. Afr. 35: 1–6.
   Google Scholar
• Bonnet, X., F. Lagarde, B.T. Henen, J, Corbin, K.A. Nagy, G. Naulleau, K. Balhoul, O. Chastel, A. Legrand & R. Cambag. 2001. Sexual dimorphism in steppe tortoises (Testudo horsfieldii): influence of the environment and sexual selection on body shape and mobility. Biol. J. Linn. SoC. 72: 357–372. doi: 10.1111/j.1095-8312.2001.tb01323.x
   Web of Science ®Google Scholar
• Bonnet, X., V. Delmas, H. El-Mouden, T. Slimani, B. Sterijovski & G. Kuchling. 2010. Is sexual body shape dimorphism consistent in aquatic and terrestrial chelonians? Zoology 113: 213–220. doi: 10.1016/j.zool.2010.03.001
   PubMed Web of Science ®Google Scholar
• Boycott, R.C. & O. Bourquin. 2000. The Southern African Tortoise Book: A Guide to Southern African Tortoises, Terrapins and Turtles. O. Borquin, Hilton, South Africa.
   Google Scholar
• Branch, W.R. 1999. Natural History Note: Homopus femoralis: Egg and clutch size. Afr. Herp News 30: 28.
   Google Scholar
• Branch, W.R. 2007. A new species of tortoise of the genus Homopus (Chelonia: Testudinidae) from southern Namibia. Afr. J. Herpetol. 56: 1–21.
   Google Scholar
• Branch, W.R. 2008. Tortoises, Terrapins and Turtles of Africa. Struik Publishers, Cape Town.
   Google Scholar
• Cooper, M.R. & D.G. Broadley. 1990. A new species of fossil Homopus (Cryptodira: Testudinidae) from South Africa. Stud. Geol. Salm. Stud. Palaeochel. 3: 41–55.
   Google Scholar
• Domokos, G. & P.L. Várkonyi. 2008. Geometry and self-righting of turtles. ProC. R. SoC. Lond. B 275: 11–17. doi: 10.1098/rspb.2007.1188
   PubMed Web of Science ®Google Scholar
• Duméril, A.M.C & G. Bibron. 1835. Erpétologie générale ou Histoire naturelle complète des Reptiles II. Paris, Roret.
   Google Scholar
• Ernst, C.H. & R.W. Barbour. 1989. Turtles of the World. Smithsonian Institution Press, Washington DC.
   Google Scholar
• Feakins, S.J. & P.B. Demenocal. 2010. Global and African Regional Climate during the CenozoiC. Pp. 45–56. In L. Werdelin & W.J. Sanders (Eds), Cenozoic Mammals of Africa. University of California Press, Berkley.
   Google Scholar
• Fitzinger, L.J. 1835. Entwurf einer svstematischen Anordnung der Schildkröten nach den Grundsätzen der natürlichen Methode. Ann. Wiener Mus. 1: 103–128.
   Google Scholar
• Fitzinger, L.J. 1843. Systema Reptilium I Amblyglossae (Conspectus geographicus). Braumueller & Seidel, Vindobonae (Wien).
   Google Scholar
• Greig, J.C. & P.D. Burdett. 1976. Patterns in the distribution of southern African terrestrial tortoises (Cryptodira: Testudinidae). Zool. Afr. 11: 249–273.
   Google Scholar
• Hewitt, J. 1931. Description of some African tortoises. Ann. Natal Mus. 6: 461–506.
   Google Scholar
• Hewitt, J. 1937. A note on the relationships of the Cape genera of land-tortoises. S. Afr. J. Sci. 33: 788–796.
   Google Scholar
• Hofmeyr, M.D., B.T. Henen & V.J.T. Loehr. 2005. Overcoming environmental and morphological constraints: egg size and pelvic kinesis in the smallest tortoise, Homopus signatus. Can. J. Zool. 83: 1343–1352. doi: 10.1139/z05-132
   Web of Science ®Google Scholar
• Hofmeyr, M.D., R.C. Boycott & E.H.W. Baard. 2014. Family Testudinidae. Pp. 70–85. In M.F. Bates, W.R. Branch, A.M. Bauer, M. Burger, J. Marais, G.J. Alexander & M.S. de Villiers (Eds), Atlas and Red List of the Reptiles of South Africa, Lesotho and Swaziland. Suricata 1. South African National Biodiversity Institute, Pretoria.
   Google Scholar
• Hofmeyr, M.D., M. Vamberger, W.R. Branch, A. Schleicher & S.R. Daniels. 2017. Tortoise (Reptilia, Testudinidae) radiations in southern Africa from the Eocene to present. Zool. Scr. 46: 389–400. doi: 10.1111/zsc.12223
   Web of Science ®Google Scholar
• Holm, S. 1979. A simple sequentially rejective multiple test procedure. Scand. J. Statist. 6: 65–70.
   Web of Science ®Google Scholar
• Keswick, T. & M.D. Hofmeyr. 2015. Sexual dimorphism and geographic variation in the morphology of a small southern African tortoise Psammobates oculifer. Amphibia-Reptilia 36: 55–64. doi: 10.1163/15685381-00002976
   Web of Science ®Google Scholar
• Le, M., C.J., Raxworthy, W.P. McCord & L. Mertz. 2006. A molecular phylogeny of tortoises (Testudines: Testudinidae) based on mitochondrial and nuclear genes. Mol. Phylogenet. Evol. 40: 517–531. doi: 10.1016/j.ympev.2006.03.003
   PubMed Web of Science ®Google Scholar
• Lindholm, W.A. 1929. Revidiertes Verzeichnis der Gattungen der rezenten Schildkröten nebst Notizen zur Nomenklatur einiger Arten. Zool. Anz. 81: 275–295.
   Google Scholar
• Loehr, V.J.T., B.T. Henen & M.D. Hofmeyr. 2004. Reproduction of the smallest tortoise, the Namaqualand speckled padloper, Homopus signatus signatus. Herpetologica 60: 444–454 doi: 10.1655/03-59
   Web of Science ®Google Scholar
• Loehr, V.J.T., B.T. Henen & M.D. Hofmeyr. 2006. Shell characteristics and sexual dimorphism in the Namaqualand speckled padloper, Homopus signatus signatus. Afr. J. Herpetol. 55: 1–11. doi: 10.1080/21564574.2006.9635536
   Web of Science ®Google Scholar
• Loehr, V.J.T. 2013. Homopus femoralis (Greater Padloper): Reproduction. Herpetol. Rev. 44: 304–305.
   Google Scholar
• Loehr, V.J.T, T. Stark, M. Weterings & H. Kuipers. 2015. Overcoming low environmental temperatures in the primary feeding season: low-level activity and long basking in the tortoise Homopus signatus. Amphibia-Reptilia 36: 207–214. doi: 10.1163/15685381-00002994
   Web of Science ®Google Scholar
• Louw. G.N. & M.K. Seely. 1982. Ecology of Desert Organisms. Longman, New York.
   Google Scholar
• Loveridge, A. & E.E. Williams. 1957. Revision of the African tortoises and turtles of the suborder Cryptodira. Bull. Mus. Comp. Zool. 115: 161–557.
   Google Scholar
• Mucina, L. & M.C. Rutherford (Eds). 2006. The Vegetation of South Africa, Lesotho and Swaziland. Strelitzia 19. South African National Biodiversity Institute, Pretoria.
   Google Scholar
• Schulze, R.E. 1997. Climate. Pp. 21–42. In R.M. Cowling, D.M. Richardson & S.M. Pierce (Eds), Vegetation of Southern Africa. Cambridge University Press, Cambridge.
   Google Scholar
• Scott, L., A. Cadman & I. McMillan. 2006. Early history of Cainozoic Asteraceae along the Southern African west coast. Rev. Palaeobot. Palynol. 142: 47–52. doi: 10.1016/j.revpalbo.2006.07.010
   Web of Science ®Google Scholar
• Turtle Taxonomy Working Group. 2017. Turtles of the World: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status (8th edn). In A.G.J. Rhodin, J.B. Iverson, P.P. van Dijk, R.A. Saumure, K.A. Buhlmann, P.C.H. Pritchard & R.A. Mittermeier (Eds). Conservation Biology of Freshwater Turtles and Tortoises: A Compilation Project of the IUCN/SSC Tortoise and Freshwater Turtle Specialist Group. Chelonian Research Monographs.
   Google Scholar
• Zug, G.R. 1968. The penial morphology and the relationships of cryptodiran turtles. Occas. Pap. Mus. Zool. Univ. MicH. 617: 1–23.
   Google Scholar