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Review Article

Hitchhiking through life: a review of phoresy in Uropodina mites (Parasitiformes: Mesostigmata)

ORCID Icon, ORCID Icon, ORCID Icon & ORCID Icon
Pages 31-63 | Received 13 Sep 2023, Accepted 23 Nov 2023, Published online: 22 Dec 2023

References

  • Ahadiyat A, Ostovan H, Saboori A. 2004. Mites associated with Scolytus amygdali Guerin-Meneville, 1847 in Karaj region. Proceeding of the 16th Iranian Plant Protection Congress, Iran, University of Tabriz. 1: Pests. pp. 251.
  • Al-Deeb MA, Muzaffar SB, Abuagla AM, Sharif EM. 2011. Distribution and abundance of phoretic mites (Astigmata, Mesostigmata) on Rhynchophorus ferrugineus (Coleoptera: Curculionidae). Florida Entomologist 94(4):748–755. DOI: 10.1653/024.094.0403.
  • Atakan E, Çobanoğlu S, Yüksel O, Bal AD. 2009. Kırmızı palmiyeböceği [Rhynchophorus ferrugineus (Oliver, 1790) (Coleoptera: Curculionidae)] üzerinde foretik uropodid akarlar (Acarina: Uropodidae) (Phoretic uropodid mites (Acarina: Uropodidae) on the red palm weevil (Rhynchophorus ferrugineus) (Olivier, 1790) (Coleoptera: Curculionidae)). Türkiye Entomoloji Dergisi 33(2):93–105.
  • Atalla EAR, Hobart J. 1964. The survival of some soil mites at different humidities and their reaction to humidity gradients. Entomologia Experimentalis et Applicata 7(3):215–228. DOI: 10.1111/j.1570-7458.1964.tb02441.x.
  • Athias F. 1976. Observations morphologiques sur Polyaspis patavinus Berlese, 1881 (Acariens: Uropodides). I. Morphologie de l’idiosoma au cours du développement postembryonnaire. Acarologia 17(3):410–435.
  • Athias-Binche F. 1981. Écologie des Uropodides édaphiques (Arachnides: Parasitifirmes) de trois écosystèmes forestiers. 1. Introduction, matériel, biologie. Vie Milieu 31(2):137–147.
  • Athias-Binche F. 1983. Acari from Operation Drake in New Guinea 2. Uropodidae. Acarologia 24(4):361–371.
  • Athias-Binche F. 1984. La phorésie chez les acariens uropodides (Anactinotriches), une stratégie écologique originale. Acta Oecologica/Oecologia Generalis 5:119–133.
  • Athias-Binche F. 1991. Evolutionary ecology of dispersal in mites. In: Dusbábek F, Bukva V, editors. Modern acarology. Vol. 1. The Hague: Academia, Prague and SPB Academic Publishing bv. pp. 27–41.
  • Athias-Binche F. 1993. Dispersal in varying environments: The case of phoretic uropodid mites. Canadian Journal of Zoology 71(9):1793–1798. DOI: 10.1139/z93-255.
  • Athias-Binche F. 1994. La phorésie chez les acariens. Aspects adaptatifs et évolutifs. Perpignan, France: Editions du Castillet. p. 178.
  • Athias-Binche F, Habersaat U. 1988. An ecological study of Janetiella pyriformis (Berlese, 1920), a phoretic Uropodina from decomposing organic matter (Acari: Anactinotrichida). Mitteilungen Der Schweizerischen Entomologischen Gesellschaft 61:377–390.
  • Athias-Binche F, Morand S. 1993. From phoresy to parasitism: The example of mites and nematodes. Research and Reviews in Parasitology 53:73–79.
  • Athias-Binche F, Schwarz HH, Meierhofer I. 1993. Phoretic association of Neoseius novus (Ouds., 1902) (Acari: Uropodina) with Nicrophorus spp. (Coleoptera: Silphidae): A case of sympatric speciation? International Journal of Acarology 19(1):75–86. DOI: 10.1080/01647959308683542.
  • Bahrami F, Arbabi M, Shoushtari RV, Kazemi S. 2011. Mesostigmatic mites associated with Coleoptera and biodiversity calculation of these mites phoretic on dung beetles in Golestan Province (North of Iran). Middle-East Journal of Scientific Research 9(3):345–366.
  • Bajerlein D. 2007. Foreza roztoczy z podrzędu Uropodina na koprofagicznych i koprofilnych chrząszczach [Phoresy of Uropodina mites on coprophagous and coprophilous beetles]. PhD- thesis. Poznań: Adam Mickiewicz University.
  • Bajerlein D. 2011. Seasonal abundance and infestation of deutonymphs of Uropoda orbicularis (Müller, 1776) (Acari: Mesostigmata) phoretic on coprophilous beetles (Scarabaeidae, Geotrupidae, Aphodiidae, Hydrophilidae, Histeridae). International Journal of Acarology 37(3):216–227. DOI: 10.1080/01647954.2010.505579.
  • Bajerlein D, Błoszyk J. 2004. Phoresy of Uropoda orbicularis by beetles (Coleoptera) associated with cattle dung in Poland. European Journal of Entomology 101(1):185–188. DOI: 10.14411/eje.2004.022.
  • Bajerlein D, Przewoźny M. 2005. Coprophagous hydrophilid beetles (Coleoptera: Hydrophilidae) as carriers of phoretic deutonymphs of Uropoda orbicularis (Acari: Mesostigmata) in Poland. European Journal of Entomology 102(1):119–122. DOI: 10.14411/eje.2005.017.
  • Bajerlein D, Przewoźny M. 2012. When a beetle is too small to carry phoretic mites? A case of hydrophilid beetles (Coleoptera: Hydrophilidae) and Uropoda orbicularis (Acari: Mesostigmata). Canadian Journal of Zoology 90(3):368–375. DOI: 10.1139/z2012-005.
  • Bajerlein D, Witaliński W. 2012. Anatomy and fine structure of pedicellar glands in phoretic deutonymphs of uropodid mites (Acari: Mesostigmata). Arthropod Structure & Development 41(3):245–257. DOI: 10.1016/j.asd.2012.02.006.
  • Bajerlein D, Witaliński W. 2014. Localization and density of phoretic deutonymphs of the mite Uropoda orbicularis (Parasitiformes: Mesostigmata) on Aphodius beetles (Aphodiidae) affect pedicel length. Naturwissenschaften 101(4):265–272. DOI: 10.1007/s00114-014-1150-x.
  • Bajerlein D, Błoszyk J, Gwiazdowicz DJ, Ptaszyk J, Halliday B. 2006. Community structure and dispersal of mites (Acari, Mesostigmata) in nests of the white stork (Ciconia ciconia). Biologia, Bratislava 61(5):525–530. DOI: 10.2478/s11756-006-0086-9.
  • Bajerlein D, Witaliński W, Adamski Z. 2013. Morphological diversity of pedicels in phoretic deutonymphs of Uropodina mites (Acari: Mesostigmata). Arthropod Structure & Development 42(3):185–196. DOI: 10.1016/j.asd.2013.02.002.
  • Bajerlein D, Adamski Z, Kacalak W, Tandecka K, Wiesner M, Jurga S. 2016. To attach or not to attach? The effect of carrier surface morphology and topography on attachment of phoretic deutonymphs of Uropoda orbicularis (Acari). The Science of Nature 103(7–8):61. DOI: 10.1007/s00114-016-1385-9.
  • Barrios-Torres PL, Villegas-Guzmán GA. 2015. Ácaros asociados a dos especies de Odontotaenius (Coleoptera: Passalidae) en México. Revista de biología tropical 63(3):659–671. DOI: 10.15517/rbt.v63i3.15745.
  • Bartlow AW, Agosta SJ. 2021. Phoresy in animals: Review and synthesis of a common but understudied mode of dispersal. Biological Reviews 96(1):223–246. DOI: 10.1111/brv.12654.
  • Barton PS, Weaver HJ, Manning AD. 2014. Contrasting diversity dynamics of phoretic mites and beetles associated with vertebrate carrion. Experimental and Applied Acarology 63(1):1–13. DOI: 10.1007/s10493-013-9758-7.
  • Beaulieu F. 2012. Saproxyly in predatory mites? Mesostigmata in decaying log habitats versus litter in a wet eucalypt forest, Tasmania, Australia. International Journal of Acarology 38(4):313–323. DOI: 10.1080/01647954.2011.647072.
  • Beaulieu F, Dowling APG, Klompen H, de Moraes GJ, Walter DE. 2011. Superorder Parasitiformes Reuter, 1909. In: Zhang Z-Q, editor. Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Vol. 3148. Auckland, New Zealand: Magnolia Press. pp. 123–128.
  • Berlese A. 1881. Indagini sulle metamorfosi di alcuni Acari insetticoli. Atti del Reale Istituto Veneto di Scienze. Lettere ed Arti Series 5(8):37–81.
  • Berlese A. 1884. Acari, Myriopoda et Scorpiones hucusque in Italia reperta,11, 13 text pages + plates 1–10. Padova: Salmin.
  • Berlese A. 1887. Acari, Myriapoda et Scorpiones hucusque in Italia reperta, 40: 13 text pages + 10 plates. Firenze: Tipographia del Seminario.
  • Berlese A. 1888a. Acari, Myriopoda et Scorpiones hucusque in Italia reperta, 49, 15 text pages + plates 1–10. Firenze: Tipographia del Seminario.
  • Berlese A. 1888b. Acari Austro-Americani quos collegit Aloysius Balzan. Manipulus primus. Species novas circiter quinquaginta complectens. Bollettino della Societa Entomologica Italiana 20:171–222. + plates V–XIII.
  • Berlese A. 1903. Acari nuovi. Manipulus I.us. Redia 1:235–252.
  • Berlese A. 1904a. Acari nuovi. Manipulus II.us. Redia 1:258–280.
  • Berlese A. 1904b. Acari nuovi. Manipulus III. Redia 2:10–32. + plates I–II.
  • Berlese A. 1904c. Illustrazione iconografica degli Acari mirmecofili. Redia 1:299–474. + plates VII–XX.
  • Berlese A. 1910. Brevi diagnosi di generi e specie novi di Acari. Redia 6:346–388.
  • Berlese A. 1916a. Centuria prima di Acari nuovi. Redia 12:19–67.
  • Berlese A. 1916b. Centuria seconda di Acari nuovi. Redia 12:125–177.
  • Berlese A. 1917. Intorno agli Uropodidae. Redia 13:7–16.
  • Berlese A. 1920. Centuria quinta di Acari nuovi. Redia 14:143–195.
  • Binns ES. 1982. Phoresy as migration - some functional aspects of phoresy in mites. Biological Reviews 57(4):571–620. DOI: 10.1111/j.1469-185X.1982.tb00374.x.
  • Błoszyk J. 1999. Geograficzne i ekologiczne zróżnicowanie zgrupowań roztoczy z kohorty Uropodina (Acari: Mesostigmata) w Polsce. I. Uropodina lasów grądowych (Carpinion betuli). Poznań: Kontekst. p. 245.
  • Błoszyk J, Bajaczyk R. 1999. The first record of Phaulodinychus borealis (Sellnick, 1940) phoresy (Acari: Uropodina) associated with fleas (Siphonaptera) in mole nests. In: Tajovský K, Pižl V, editors. Soil Zoology in Central Europe; Proceedings of the 5th Central European Workshop on Soil Zoology. České Budějovice, Czech Republic: Institute of Soil Biology, Academy of Sciences of the Czech Republic. pp. 13–17.
  • Błoszyk J, Adamski Z, Napierała A, Dylewska M. 2004. Parthenogenesis as a life strategy among mites of the suborder Uropodina (Acari: Mesostigmata). Canadian Journal of Zoology 82(9):1503–1511. DOI: 10.1139/z04-133.
  • Błoszyk J, Klimczak J, Leśniewska M. 2006. Phoretic relationships between Uropodina (Acari: Mesostigmata) and centipedes (Chilopoda) as an example of evolutionary adaptation of mites to temporary microhabitats. European Journal of Entomology 103(3):699–707. DOI: 10.14411/eje.2006.093.
  • Błoszyk J, Gutowski JM, Gwiazdowicz DJ, Mądra A. 2013. Phoresy of Trichouropoda shcherbakae Hirschmann, 1972 (Acari: Mesostigmata) on beetles of the genus Tetropium Kirby, 1837 (Coleoptera: Cerambycidae) in Białowieża forest, Poland. Central European Journal of Biology 8(10):986–992. DOI: 10.2478/s11535-013-0226-5.
  • Błoszyk J, Gutowski JM, Gwiazdowicz DJ, Mądra A, Konwerski S, Książkiewicz Z. 2016. Monochamus sartor (Coleoptera: Cerambycidae) contributes to alpha diversity of Uropodina mites (Acari: Mesostigmata) in first stage of wood decay in Białowieża Primeval forest. International Journal of Acarology 42(4):218–223. DOI: 10.1080/01647954.2016.1154890.
  • Brückner A, Klompen H, Bruce AI, Hashim R, von Beeren C. 2017. Infection of army ant pupae by two new parasitoid mites (Mesostigmata: Uropodina). PeerJ 5:e3870. DOI: 10.7717/peerj.3870.
  • Cai C, Tihelka E, Giacomelli M, Lawrence JF, Ślipiński A, Kundrata R, Yamamoto S, Thayer MK, Newton AF, Leschen RAB, Gimmel ML, Lü L, Engel MS, Bouchard P, Huang D, Pisani D, Donoghue PCJ. 2022. Integrated phylogenomics and fossil data illuminate the evolution of beetles. Royal Society Open Science 9(3):211771. DOI: 10.1098/rsos.211771.
  • Camerik AM. 2010. Phoresy revisited. In: Sabelis M, Bruin J, editors. Trends in Acarology, Proceedings of the 12th International Congress of Acarology, Amsterdam, The Netherlands, Dordrecht Heidelberg London New York, Springer Science+Business Media B.V. pp. 333–336.
  • Campbell KU, Klompen H, Crist TO. 2013. The diversity and host specificity of mites associated with ants: The roles of ecological and life history traits of ant hosts. Insectes Sociaux 60(1):31–41. DOI: 10.1007/s00040-012-0262-6.
  • Canestrini G, Berlese A. 1884. Sopra alcune nuove specie di Acari Italiani. Atti della Societá Veneto-Trentina di Scienze Naturali 9:175–182 + plates 3–5.
  • Čejka M, Holuša J. 2014a. Phoretic mites in uni - and bivoltine populations of Ips typographus: A 1-year case study. Turkish Journal of Zoology 38(5):569–574. DOI: 10.3906/zoo-1309-20.
  • Čejka M, Holuša J. 2014b. Foretičtí roztoči lýkožrouta severského Ips duplicatus (Coleoptera: Curculionidae: Scolytinae) z oblasti recentní gradace ve střední Evropě. Lesnicky Časopis – Forestry Journal 60(4):240–243. DOI: 10.1515/forj-2015-0006.
  • Chaires-Grijalva MP, Estrada-Venegas EG, Equihua-Martínez A, Moser JC, Sánchez-Martínez G, Vásquez-Rojas IM, Otero-Colina G, Romero-Nápoles J. 2013. Mesostigmados (Acari: Mesostigmata) asociados con Dendroctonus rhizophagus de Chihuahua, México [Mesostigmatid mites (Acari, Mesostigmata) associated to Dendroctonus rhizophagus from Chihuahua, Mexico]. Revista Mexicana De Biodiversidad 84(4):1235–1242. DOI: 10.7550/rmb.35723.
  • Chaires-Grijalva MP, Estrada-Venegas EG, Equihua-Martinez A, Moser JC, Blomquist SR. 2016. Trophic habits of mesostigmatid mites associated with bark beetles in Mexico. Journal of the Acarological Society of Japan 25(S1):161–167. DOI: 10.2300/acari.25.Suppl_161.
  • Chmielewski W. 1997 1998. Roztocze (Acarina) występujące na pszczołach społecznych (Hymenoptera: Apidae: Apinae, Bombinae). [Mites (Acarina) occurring on social bees (Hymenoptera: Apidae: Apinae, Bombinae)]. Wiadomości Entomologiczne 16(3–4):201–216.
  • Chmielewski W. 2000. Powiązania roztoczy z chrząszczami (Acarina-Coleoptera (Insecta)). In: Ignatowicz S, editor. Akarologia polska u progu XXI wieku. Proceedings of the XXVI Acarological Syposium, Kazimierz Dolny, 24-26 October 1999, Warszawa, Poland. pp. 79–90.
  • Cilbircioğlu C, Kovač M, Pernek M. 2021. Associations of phoretic mites on bark beetles of the genus Ips in the Black Sea Mountains of Turkey. Forests 12(5):516. DOI: 10.3390/f12050516.
  • Constantinescu IC, Cristescu C. 2007. Studies on the trophic preferences of certain Uropodina mites (Acarina: Anactinotrichida, Uropodina) for some taxa of imperfect fungi. Analele Ştiinţifice ale Universităţii ,Al I Cuza” Iaşi, s Biologie animală 53:83–87.
  • Cooreman J. 1963. Notes et observations sur quelques Acariens inféodés aux Coléoptères Scolytides de la faune Belge. Bulletin de l’Institut Royal des Sciences Naturelles de Belgique 39:1–48.
  • Costa M. 1963. The mesostigmatic mites associated with Copris hispanus (L.) (Coleoptera, Scarabaeidae) in Israel. Journal of the Linnaean Society of London, Zoology 45(303):25–45. DOI: 10.1111/j.1096-3642.1963.tb00485.x.
  • Costa M. 1966. The present stage of knowledge of mesostigmatic mites in Israel (Acari, Mesostigmata). Israel Journal of Zoology 15(2):69–82.
  • Danilevsky M. 2020. Catalogue of Palaearctic Coleoptera. Volume 6/1. Chrysomeloidea I (Vesperidae, Disteniidae, Cerambycidae). Updated and revised second. Leiden, Boston: Brill. xxvii +. p. 712.
  • De Geer C. 1768. Djur, som med en sträng i ändan äro säste vid andra lesvande djur [Animals that, with a string at the end, are attached to other living animals]. Kongliga Vetenskaps Academiens Handlingar 29:176–183 + Plate IV.
  • Delfinado-Baker M, Rath W, Boecking O. 1992. Phoretic bee mites and honeybee grooming behavior. International Journal of Acarology 18(4):315–322. DOI: 10.1080/01647959208683966.
  • Desender K, Vaneechoutte M. 1984. Phoretic associations of carabid beetles (Coleoptera, Carabidae) and mites (Acari). Revue d’Ecologie et Biologie du Sol 21:363–371.
  • Dilipkumar M, Ahadiyat A, Mašán P, Chuah TS. 2015. Mites (Acari) associated with Rhynchophorus ferrugineus (Coleoptera: Curculionidae) in Malaysia, with a revised list of the mites found on this weevil. Journal of Asia-Pacific Entomology 18(2):169–174. DOI: 10.1016/j.aspen.2014.12.010.
  • Domrow R. 1981. A small lizard stifled by phoretic deutonymphal mites (Uropodina). Acarologia 22(3):247–252.
  • Dunlop JA, Kontschán J, Zwanzig M. 2013. Fossil mesostigmatid mites (Mesostigmata: Gamasina, Microgyniina, Uropodina) associated with longhorn beetles (Coleoptera: Cerambycidae) in Baltic amber. Naturwissenschaften 100(4):337–344. DOI: 10.1007/s00114-013-1031-8.
  • Eickwort GC. 1990. Associations of mites with social insects. Annual Review of Entomology 35(1):469–488. DOI: 10.1146/annurev.en.35.010190.002345.
  • El-Banhawy EM, El-Borolossy MA, El-Sawaf BM, Afia SI. 1997. Biological aspects and feeding behaviour of the predacious soil mite Nenteria hypotrichus (Uropodina: Uropodidae). Acarologia 38(4):357–360.
  • Elzinga RJ. 1978. Holdfast mechanisms in certain Uropodine mites (Acarina: Uropodina). Annals of the Entomological Society of America 71(6):896–900. DOI: 10.1093/aesa/71.6.896.
  • Faasch H. 1967. Beitrag zur Biologie der einheimischen Uropodiden Uroobovella marginata (C. L. Koch 1839) und Uropoda orbicularis (O. F. Müller 1776) und experimentelle Analyse ihres Phoresieverhaltens. Zoologische Jahrbücher, Abteilung für Systematik 94:521–608.
  • Fain A. 1998a. Description of mites (Acari) phoretic on Phoridae (Insecta: Diptera) with description of four new species of the genus Uroseius Berlese (Parasitiformes, Uropodina, Polyaspididae). International Journal of Acarology 24(3):213–220. DOI: 10.1080/01647959808683586.
  • Fain A. 1998b. New mites (Acari) phoretic on Phoridae and Ephydridae (Diptera) from Thailand. Bulletin de L’institut Royal Des Sciences Naturelles de Belgique Entomologie 68:53–61.
  • Fain A, Greenwood MT. 1991. Notes on a small collection of mites (Acari), phoretic on Diptera, mainly Phoridae, from the British Isles. Bulletin de L’institut Royal Des Sciences Naturelles de Belgique Entomologie 61:193–197.
  • Farish JD, Axtell RC. 1971. Phoresy redefined and examined in Macrocheles muscaedomesticae (Acarina: Macrochelidae). Acarologia 13(1):16–29.
  • Farmahiny Farahani VR, Ahadiyat A, Mašán P, Dehvari MA. 2016. Phoretic uropodine mites (Acari: Mesostigmata) associated with the red palm weevil, Rhynchophorus ferrugineus (Coleoptera: Curculionidae) in Iran. Journal of Entomological and Acarological Research 48(3):5853. DOI: 10.4081/jear.2016.5853.
  • Feketeová Z. 2011a. Lykožrút smrekový (Ips typographus) a jeho úloha vo forézii uropodných roztočov (Acari: Uropodina). Entomofauna carpathica 23(1):23–30.
  • Feketeová Z. 2011b. Význam Uropodných roztočov (Acari: Uropodina) v ekológii lykožrúta smrekového (Ips typographus). Entomofauna carpathica 23(2):11–19.
  • Fernández M, Diez J, Moraza ML. 2013. Acarofauna associated with Ips sexdentatus in northwest Spain. Scandinavian Journal of Forest Research 28(4):358–362. DOI: 10.1080/02827581.2012.745897.
  • Goff ML. 1989. Gamasid mites as potential indicators of postmortem interval. In: Channabasavanna G, Viraktamath C, editors. Progress in acarology-volume 1. New Delhi, India: Oxford and IBH. pp. 443–450.
  • Goff ML. 1991. Use of Acari in establishing a postmortem interval in a homicide case on the island of Oahu, Hawaii. In: Dusbábek F, Bukva V, editors. Modern acarology. Academia, Prague and SPB. Vol. 1. The Hague, Netherlands: Academic Publishing bv. pp. 439–442.
  • Gómez-Marco F, Klompen H, Hoddle MS. 2021. Phoretic mite infestations associated with Rhynchophorus palmarum (Coleoptera: Curculionidae) in Southern California. Systematic & Applied Acarology 26(10):1913–1926. DOI: 10.11158/saa.26.10.6.
  • Gordh G. 1985. Uropoda sp. phoretic on Elater lecontei Horn. Pan - Pacific Entomologist 6:154.
  • Gordh G, Barrows EM. 1976. Uropoda phoretic on Ceratina (Acarina: Hymenoptera). Journal of the Kansas Entomological Society 49(3):344–345.
  • Gordh G, Wills L. 1989. Anatomical notes on Uropoda sp., a phoretic mite infesting dung- inhabiting beetles in Southern California (Acari: Uropodidae; Coleoptera: Tenebrionidae, Histeridae). Pan-Pacific Entomologist 65(4):410–413.
  • Gwiazdowicz DJ. 2000. Mites (Acari: Gamasida) associated with insects in the Białowieża National Park. Acta Parasitologica 45(1):43–47. DOI: 10.3161/00159301FF2000.43.8.091.
  • Gwiazdowicz DJ, Gutowski JM. 2012. Records of phoretic mesostigmatid mites (Acari: Mesostigmata) on beetles (Coleoptera: Carabidae, Cerambycidae, Elateridae, Erotylidae, Scolytinae) in the Białowieża Primeval forest. Polish Journal of Entomology 81(4):305–310. DOI: 10.2478/v10200-012-0010-2.
  • Gwiazdowicz DJ, Kamczyc J, Błoszyk J. 2011. The diversity of phoretic Mesostigmata on Ips typographus (Coleoptera: Scolytinae) caught in the Karkonosze forest. European Journal of Entomology 108(3):489–491. DOI: 10.14411/eje.2011.063.
  • Gwiazdowicz DJ, Kamczyc J, Teodorowicz E, Błoszyk J. 2012. Mite communities (Acari, Mesostigmata) associated with Ips typographus (Coleoptera, Scolytidae) in managed and natural Norway spruce stands in Central Europe. Central European Journal of Biology 7:910–916. DOI: 10.2478/s11535-012-0070-z.
  • Gwiazdowicz DJ, Gutowski JM, Kamczyc J, Teodorowicz E. 2013. Phoretic relationships between Plagionotus detritus (Coleoptera: Cerambycidae) and Trichouropoda sociata (Acari: Mesostigmata). Entomologica Fennica 24(1):59–64. DOI: 10.33338/ef.7868.
  • Gwiazdowicz DJ, Błoszyk J, Gdula AK. 2015. Alpha diversity of mesostigmatid mites associated with the bark beetle Ips typographus (L.) in Poland. Insect Conservation and Diversity 8(5):448–455. DOI: 10.1111/icad.12122.
  • Halliday RB. 2015. Catalogue of genera and their type species in the mite suborder Uropodina (Acari: Mesostigmata). Zootaxa 3972(2):101–147. DOI: 10.11646/zootaxa.3972.2.1.
  • Halliday RB. 2016. Catalogue of families and their type genera in the mite suborder Uropodina (Acari: Mesostigmata). Zootaxa 4061(4):347–366. DOI: 10.11646/zootaxa.4061.4.2.
  • Halliday B. 2019. The enemy of my parasite is my friend: The possible role of predatory mites as biological control agents of pest beetles in soil. International Journal of Acarology 45(4):189–196. DOI: 10.1080/01647954.2019.1574895.
  • Helson GAH, Flower NE, Wood J. 1975. Phoretic association of a uropodid mite and a lepidopteran larva. New Zealand Journal of Zoology 2(2):223–226. DOI: 10.1080/03014223.1975.9517872.
  • Hirschmann W. 1972a. Gangsystematik der Parasitiformes. Teil 92. Gänge, Teilgänge, Stadien von 13 neuen Trichouropoda-Arten (Trichouropodini, Uropodinae). Acarologie Schriftenreihe für Vergleichende Milbenkunde 17:3–8 + plates 1–3.
  • Hirschmann W. 1972b. Gangsystematik der Parasitiformes. Teil 93. Gänge, Teilgang, Stadien von 7 neuen Uroobovella-Arten (Dinychini, Uropodinae). Acarologie Schriftenreihe für Vergleichende Milbenkunde 17:9–13 + plates 3–4.
  • Hirschmann W. 1975. Gangsystematik der Parasitiformes. Teil 193. Teilgang einer neuen Trichouropoda-Art (Trichouropodini, Uropodinae). Acarologie Schriftenreihe für Vergleichende Milbenkunde 21:10–12. + plate 9.
  • Hirschmann W. 1979. Stadiensystematik der Parasitiformes. Teil 1. Stadienfamilien und Stadiengattungen der Atrichopygidiina, erstellt im Vergleich zum Gangsystem Hirschmann 1979. Acarologie Schriftenreihe für Vergleichende Milbenkunde 26:57–70.
  • Hirschmann W. 1981. Gangsystematik der Parasitiformes. Teil 420. Teilgang und Stadien von 4 neuen Uroobovella-Arten der Pulchella- und Carinata-Gruppe aus Belgien, Zaire, USA und Vietnam (Dinychini, Uropodinae) Acarologie. Schiftenreihe für vergleichende Milbenkunde 28:125–127.
  • Hirschmann W. 1985. Gangsystematik der Parasitiformes. Teil 489. Stadien von 2 neuen Nenteria-Arten der rühmi-Gruppe an Pyragra (Dermaptera) aus Argentinien und Bolivien (Trichouropodini, Uropodinae) Acarologie. Schiftenreihe für vergleichende Milbenkunde 32:163–166.
  • Hirschmann W. 1989. Gangsystematik der Parasitiformes. Teil 509. Die Ganggattung Uroobovella Berlese 1903 – Artengruppen – Bestimmungstabellen – Diagnosen – (Dinychini, Uropodinae). Acarologie Schiftenreihe für vergleichende Milbenkunde 36:84–196.
  • Hirschmann W, Kemnitzer F. 1989. Polyaspis (Polyaspis)-Deutonymphen Polyaspis-Wandernymphentraube an Solitärwespe (Scoliidae) (Atrichopygidiina: Uropodina). Acarologia 30(1):3–12.
  • Hirschmann W, Wiśniewski J. 1985. Gangsystematik der Parasitiformes. Teil 486. Die rühmi-Gruppe, eine neue Adulten-Gruppe der Ganggattung Nenteria Gang Teilgänge, Stadien von 12 neuen Nenteria-Arten aus Israel, Venezuela, Ekuador, Guatemala, Tanzania, Rwanda, Java, Sumatra, Laos, Neuguinea, Neu-Britannien. Neuzeichnungen und Ergänzungsbeschreibungen von 3 bekannten Nenteria-Arten (Trichouropodini, Uropodinae) Acarologie Schiftenreihe für vergleichende Milbenkunde 32:96–132.
  • Hirschmann W, Wiśniewski J. 1986. Gangsystematik der Parasitiformes. Teil 495. Weltweite Revision der Ganggattung Trichouropoda Berlese 1916. V. Die obscura-Gruppe (Trichouropodini, Uropodinae). Acarologie Schriftenreihe für Vergleichende Milbenkunde 33:149–176.
  • Hirschmann W, Wiśniewski J. 1987. Gangsystematik der Parasitiformes. Teil 497. Weltweite Revision der Ganggattung Trichouropoda Berlese, 1916 VII. Die sociata-Gruppe (Trichouropodini, Uropodinae). Acarologie Schiftenreihe für vergleichende Milbenkunde 34:51–132.
  • Hirschmann W, Wiśniewski J. 1989. Gangsystematik der Parasitiformes. Teil 507-508. Weltweite Revision der Gattung Trichouropoda Berlese 1916 Teil IV. Acarologie Schiftenreihe für vergleichende Milbenkunde 36:1–72.
  • Hirschmann W, Zirngiebl-Nicol I. 1961. Gangsystematik der Parasitiformes. Teil 4. Die Gattung Trichouropoda Berlese 1916 nov. comb., die Cheliceren und das System der Uropodiden. Acarologie Schriftenreihe für Vergleichende Milbenkunde 4:1–41 + plates 1–16.
  • Hirschmann W, Zirngiebl-Nicol I. 1969. Gangystematik der Parasitiformes. Teil 73. Neuzeichung bekannter Uropodiden-Arten, Gänge, Teilgänge, Stadien, Chaetotaxie, Literatur, Synonyma, Fundorte, Grösse. Acarologie Schiftenreihe für vergleichende Milbenkunde 12:125–132.
  • Hofstetter RW, Moser JC, Blomquist SR. 2013. Mites associated with bark beetles and their hyperphoretic ophiostomatoid fungi. In: Seifert K, De Beer Z, Wingfield M, editors. The ophiostomatoid fungi: Expanding frontiers. Vol. 12. Utrecht: CBS-KNAW Fungal Biodiversity Centre. pp. 165–176.
  • Holuša J, Čejka M. 2020. Estimates of phoretic mite abundance on bark beetles as affected by beetle capture method: A case study with Mesostigmata mites and Ips typographus (Coleoptera: Curculionidae). Experimental and Applied Acarology 82(3):347–357. DOI: 10.1007/s10493-020-00560-9.
  • Houck MA, OConnor BM. 1991. Ecological and evolutionary significance of phoresy in the Astigmata. Annual Reviews in Entomology 36(1):611–636. DOI: 10.1146/annurev.en.36.010191.003143.
  • Hunter PE, Rosario RMT. 1988. Associations of Mesostigmata with other arthropods. Annual Review of Entomology 33(1):393–417. DOI: 10.1146/annurev.en.33.010188.002141.
  • Hyatt KH. 1959. Mesostigmatic mites associated with Geotrupes stercorarius (L.) (Col., Scarabaeidae). Entomologist’s Monthly Magazine 95:22–23.
  • Ibrahim M, Loulou A, Brouk A, Muller A, Machado RAR, Kallel S. 2021. Parasites rather than phoronts: Teratorhabditis synpapillata nematodes reduce lifespan of their Rhynchophorus ferrugineus host in a life stage-dependent manner. Ecology and Evolution 11(18):12596–12604. DOI: 10.1002/ece3.8004.
  • Iwan D, Löbl I. 2020. Catalogue of Palaearctic Coleoptera. Volume 5. Tenebrionoidea. Revised and updated second. Leiden, Boston: Brill xxiv +. p. 945.
  • Jałoszyński P. 2015. Ptenidium pusillum (Gyllenhal, 1808) from egg to pupa (Coleoptera: Ptiliidae). Zootaxa 3948(3):361–421. DOI: 10.11646/zootaxa.3948.3.3.
  • Kamczyc J, Gwiazdowicz DJ. 2013. The diversity of soil mites (Acari: Mesostigmata) in yellow ant (Lasius flavus) nests along a gradient of land use. Biologia 68(2):314–318. DOI: 10.2478/s11756-013-0154-x.
  • Kapuściński S, Wiśniewski J. 1985. Wandernymphen von Milben als Passagiere auf Käfern. Mikrokosmos 74(6):161–163.
  • Karg W. 1989. Acari (Acarina), Milben. Unterordnung Parasitiformes (Anactinochaeta). Uropodina Kramer, Schlidkrötenmilben. Die Tierwelt Deutschlands 67:1–203.
  • Khaustov AA, Trach VA, Bobylev AN. 2016. Mites (Acari) phoretic on six-toothed spruce bark beetle, Pityogenes chalcographus Linnaeus (Coleoptera: Curculionidae: Scolytinae), in western Siberia, Russia. Acarina 24(2):137–151. DOI: 10.21684/0132-8077-2016-24-2-137-151.
  • Khaustov AA, Klimov PB, Trach VA, Bobylev AN, Salavatulin VM, Khaustov VA, Tolstikov AV. 2018. Review of mites (Acari) associated with the European spruce bark beetle, Ips typographus (Coleoptera: Curculionidae: Scolytinae) in Asian Russia. Acarina 26(1):3–79. DOI: 10.21684/0132-8077-2018-26-1-3-79.
  • Kinn DN. 1982. Seasonal distribution of three common mite associates of the southern pine beetle (Coleoptera: Scolytidae) in central Louisiana. The Florida Entomologist 65(1):185–187. DOI: 10.2307/3494161.
  • Kinn DN, Linit MJ. 1989. A key to phoretic mites commonly found on long-horned beetles emerging from Southern Pines. New Orleans, LA, USA: United States Department of Agriculture, Forest Service Southern Forest Experiment Station. Research Note SO-357. pp. 1–8. DOI: 10.2737/SO-RN-357.
  • Kinn DN, Witcosky JJ. 1978. Variation in southern pine beetle attack height associated with phoretic uropodid mites. The Canadian Entomologist 110(3):249–251. DOI: 10.4039/Ent110249-3.
  • Klausnitzer B. 2018. Schildkrötenmilben (Uropodina) an Calvia decemguttata (Linnaeus, 1767) (Coleoptera, Coccinellidae). Entomologische Nachrichten und Berichte 62:71–72.
  • Knee W, Beaulieu F, Skevington JH, Kelso S, Cognato AI, Forbes MR, Shaw P. 2012a. Species boundaries and host range of tortoise mites (Uropodoidea) phoretic on bark beetles (Scolytinae), using morphometric and molecular markers. PloS One 7(10):e47243. DOI: 10.1371/journal.pone.0047243.
  • Knee W, Beaulieu F, Skevington JH, Kelso S, Forbes MR. 2012b. Cryptic species of mites (Uropodoidea: Uroobovella spp.) associated with burying beetles (Silphidae: Nicrophorus): The collapse of a host generalist revealed by molecular and morphological analyses. Molecular and Phylogenetics and Evolution 65(1):276–286. DOI: 10.1016/j.ympev.2012.06.013.
  • Knee W, Forbes MR, Beaulieu F. 2013. Diversity and host use of mites (Acari: Mesostigmata, Oribatida) phoretic on bark beetles (Coleoptera: Scolytinae): Global generalists, local specialists? Annals of the Entomological Society of America 106(3):339–350. DOI: 10.1603/AN12092.
  • Knee W, Hartzenberg T, Forbes MR, Beaulieu F. 2012c. The natural history of mites (Acari: Mesostigmata) associated with the white-spotted sawyer beetle (Coleoptera: Cerambycidae): Diversity, phenology, host attachment, and sex bias. The Canadian Entomologist 144(5):711–719. DOI: 10.4039/tce.2012.57.
  • Kočárek P. 2003. Decomposition and Coleoptera succession on exposed carrion of small mammal in Opava, the Czech Republic. European Journal of Soil Biology 39(1):31–45. DOI: 10.1016/S1164-5563(02)00007-9.
  • Koch CL. 1835. Deutschlands Crustaceen, Myriapoden und Arachniden. Ein Beitrag zur Deutschen Fauna. G.A.W Herrich‐Schäffer, editor. Regensberg: Putset. 2:25 un-numbered text pages + 24 plates.
  • Koch CL. 1840. Deutschlands Crustaceen, Myriapoden und Arachniden. Ein Beitrag zur Deutschen Fauna. G.A.W Herrich‐Schäffer, editor. Regensberg: Montag & Weiβ. 27:25 un-numbered text pages + 24 plates.
  • Kofler A, Schmölzer K. 2000. Zur Kenntnis phoretischer Milben und ihrer Tragwirte in Österreich (Acarina: Gamasina: Uropodina). Berichte des Naturwissenschaftlich-medizinischen Vereins in Innsbruck 87:133–157.
  • Kontschán J, Tambe JT, Riolo P. 2012. Uroobovella phoenicicola sp. n., a new Uropodina mite (Acari: Mesostigmata) associated with the African palm weevil (Rhynchophorus phoenicis Fabricius, 1801) from Cameroon. African Invertebrates 53(2):593–600. DOI: 10.5733/afin.053.0205.
  • Kontschán J, Mazza G, Nannelli R, Roversi PF. 2014a. Nenteria extremica n. sp., a new Uropodina mites (Acari Mesostigmata) collected on Rhynchophorus ferrugineus in Italy, with notes on other Uropodina mites associated with red palm Weevil. Redia 97:63–69.
  • Kontschán J, Mazza G, Nannelli R, Roversi PF. 2014b. The true identity of the red palm weevil associated Uropodina mite, Centrouropoda almerodai Hiramatsu & Hirschmann. Redia 97:83–88.
  • Konwerski S, Gutowski JM, Książkiewicz-Parulska Z, Błoszyk J. 2017. Repeatability of the phoretic relationships between mites of the genus Trichouropoda Berlese (Parasitiformes: Uropodina) and longhorn beetles of the genus Tetropium Kirby (Coleoptera: Cerambycidae) in Białowieża Primeval forest, Central Europe. International Journal of Acarology 43(8):612–621. DOI: 10.1080/01647954.2017.1379558.
  • Konwerski S, Gutowski JM, Błoszyk J. 2019. Analysis of the phoretic relationships between mites of the genus Trichouropoda Berlese (Parasitiformes: Uropodina) and the longhorn beetle Plagionotus detritus (Linnaeus) (Coleoptera: Cerambycidae) based on multiannual observations in Białowieża Primeval forest, Central Europe. International Journal of Acarology 45(1–2):29–40. DOI: 10.1080/01647954.2018.1538257.
  • Konwerski S, Gutowski JM, Błoszyk J. 2020. Patterns of distribution of phoretic deutonymphs of Uropodina on longhorn beetles in Białowieża Primeval Forest, Central Europe. Diversity 12(6):239. DOI: 10.3390/d12060239.
  • Kotiaho JS, Simmons LW. 2001. Effects of Macrocheles mites on longevity of males of the dimorphic dung beetle Onthophagus binodis. Journal of Zoology (London, England) 254(4):441–445. DOI: 10.1017/S0952836901000930.
  • Kramer P. 1886. Ueber Milben. Archiv für Naturgeschichte 52:241–268. + plate XII. DOI: 10.5962/bhl.part.28438.
  • Krištofík J, Mašán P, Šustek Z, Gajdoš P. 1993. Arthropods in the nests of penduline tit (Remiz pendulinus). Biologia, Bratislava 48(5):493–505.
  • Kršiak B, Zach P, Kulfan J. 2010. The role of Hylastes cunicularius Erichson (Coleoptera: Scolytidae) in transferring uropodine mites in a mountain spruce forest. Journal of Forest Science 56(6):258–264. DOI: 10.17221/81/2009-JFS.
  • Kuhlmann M. 1998. Nachweise mit Bienen und Wespen (Hymenoptera Aculeata) assoziierter Milben (Acari) und Fächerflügler (Strepsiptera). Linzer Biologische Beiträge 30(1):69–80.
  • Lachaud J-P, Klompen H, Pérez-Lachaud G. 2016. Macrodinychus mites as parasitoids of invasive ants: An overlooked parasitic association. Scientific Reports 6:29995. DOI: 10.1038/srep29995.
  • Latreille PA. 1806. Genera Crustaceorum et Insectorum secundum ordinem naturalem in famlias disposita, iconibus exemplisque plurimus explicita. Vol. 1. König, Paris. xviii + 302. p. + 16 plates. DOI: 10.5962/bhl.title.65741.
  • Lehtinen PT. 1987. Association of uropodid, prodinychid, polyaspidid, antennophorid, sejid, microgynid, and zerconid mites with ants. Entomologisk Tidskrift 108:13–20.
  • Levieux J, Lieutier F, Moser JC, Perry TJ. 1989. Transportation of phytopathogenic fungi by the bark beetle Ips sexdentatus Boerner and associated mites. Journal of Applied Entomology 108(1–5):1–11. DOI: 10.1111/j.1439-0418.1989.tb00425.x.
  • Lieutier F. 1978. Les acariens associés a Ips typographus et Ips sedentatus (Coleoptera: Scolytidae) en région parisienne et les variations de leurs populations au cours du cycle annuel. Bulletin de la Société d’Ecologie 9(4):307–321.
  • Löbl I, Löbl D. 2015. Catalogue of Palaearctic Coleoptera. Hydrophiloidea – Staphylinoidea. Volume 2. Revised and updated ed. Leiden, Boston: Brill. xxvi +. p. 1702.
  • Löbl I, Löbl D. 2016. Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea – Scirtoidea – Dascilloidea – Buprestoidea – Byrrhoidea. Revised and updated ed. Leiden, Boston: Brill. xxviii +. p. 983.
  • Löbl I, Löbl D. 2017. Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata – Myxophaga – Adephaga. Revised and updated ed. Leiden, Boston: Brill. xxxiv +. p. 1443.
  • Löbl I, Smetana A. 2007. Catalogue of Palaearctic Coleoptera. Volume 4. Elateroidea, Derodontoidea, Bostrichoidea, Lymexyloidea, Cleroidea, Cucujoidea. Stenstrup: Apollo Books. p. 935.
  • Löbl I, Smetana A. 2010. Catalogue of Palaearctic Coleoptera. Volume 6. Chrysomeloidea. Stenstrup: Apollo Books. p. 924.
  • Löbl I, Smetana A. 2011. Catalogue of Palaearctic Coleoptera. Volume 7. Curculionoidea I. Stenstrup: Apollo Books. pp. 373.
  • Löbl I, Smetana A. 2013. Catalogue of Palaearctic Coleoptera. Volume 8. Curculionoidea II. Leiden: Brill. p. 707.
  • Longo S, Ragusa Di Chiara S. 2006. Presenza e diffusione in Italia dell’acaro Centrouropoda almerodai (Uroactiniinae, Uropodina). Bollettino di Zoologia Agraria e di Bachicoltura 38(3):265–269.
  • Longo S, Suma P, Bella S, La Pergola A. 2009. Artropodi associate al Punteruolo rosso delle palme. Regione Siciliana—Assessorato Agricoltura e Foreste Dipartimento Interventi Infrastrutturali—Servizi allo Sviluppo, editor La ricerca scientifica sul Punteruolo rosso e gli altri fitofagi delle palme in Sicilia 1:61–64.
  • Loulou A, Fakhraoui A, M’rabet‑Samaali B, Chihani‑Hammas N, Kallel S. 2023. Interactions between microorganisms associated with red palm weevil, Rhynchophorus ferrugineus, in Tunisia. Euro-Mediterranean Journal for Environmental Integration 8(3):493–505. DOI: 10.1007/s41207-023-00382-2.
  • Majka CG, Behan-Pelletier V, Bajerlein D, Błoszyk J, Krantz GW, Lucas Z, OConnor B, Smith IM. 2007. New records of mites (Arachnida: Acari) from Sable Island, Nova Scotia, Canada. The Canadian Entomologist 139(5):690–699. DOI: 10.4039/n06-103.
  • Makarova OL. 1995. Mesostigmatic mites (Parasitiformes, Mesostigmata) on the forest dung beetle Geotrupes stercorosus. Zoologicheskii Zhurnal. 74(12):16–23.
  • Mašán P. 1993. Mites (Acarina) associated with species of Trox (Coleoptera: Scarabaeidae). European Journal of Entomology 90:359–364.
  • Mašán P. 1994. The mesostigmatic mites (Acarina, Mesostigmata) associated with the dung beetles (Coleoptera, Scarabaeidae) in South Slovakia. Biologia, Bratislava 49:201–205.
  • Mašán P. 1998. Description of the female of Uropoda copridis (Acarina, Mesostigmata, Uropodina). Biologia, Bratislava 53:651–653.
  • Măsán P. 1999. New mite species of the cohort Uropodina (Acarina, Mesostigmata) from Slovakia. Biologia, Bratislava 54(2):121–133.
  • Mašán P. 2001. Mites of the cohort Uropodina (Acarina, Mesostigmata) in Slovakia. Annotationes Zoologicae et Botanicae 223:1–321.
  • Mašán P, Halliday B. 2009. Mesostigmatid mites associated with the dung beetle Copris lunaris (Coleoptera: Scarabaeidae). European Journal of Entomology 106(4):545–550. DOI: 10.14411/eje.2009.068.
  • Matos I, Silva D, Oliveira J, Gonçalves C, Alves R, Pereira N, Catarino F, Ameixa OMCC, Sousa JA, Rangel LF, Santos MJ, Ayra-Pardo C. 2023. Body size-dependent effects on the distribution patterns of phoretic mite species assemblages on Rhynchophorus ferrugineus (Olivier, 1790). Ecology and Evolution 13(7):e10338. DOI: 10.1002/ece3.10338.
  • Mazza G, Cini A, Cervo R, Longo S. 2011. Just phoresy? Reduced lifespan in red palm weevils Rhynchophorus ferrugineus (Coleoptera: Curculionidae) infested by the mite Centrouropoda almerodai (Uroactiniinae: Uropodina). Italian Journal of Zoology 78(1):101–105. DOI: 10.1080/11250003.2010.509135.
  • Mazza G, Francardi V, Simoni S, Benvenuti C, Cervo R, Faleiro JR, Llácer E, Longo S, Nannelli R, Tarasco E, Roversi PF. 2014. An overview on the natural enemies of Rhynchophorus palm weevils, with focus on R ferrugineus. Biological Control 77:83–92. DOI: 10.1016/j.biocontrol.2014.06.010.
  • McGarry JW, Baker AS. 1997. Observations on the mite fauna associated with adult Stomoxys calcitrans in the U.K. Medical and Veterinary Entomology 11(2):159–164. DOI: 10.1111/j.1365-2915.1997.tb00307.x.
  • Mertins JW, Hartdegen RW. 2003. The ground skink, Scincella lateralis, an unusual host for phoretic deutonymphs of a uropodine mite, Fuscuropoda marginata with a review of analogous mite-host interactions. The Texas Journal of Science 55(1):33–42.
  • Michael AD. 1894. Notes on the Uropodinæ. Journal of the Royal Microscopical Society 1894(3):289–319 + plates VI–VII. DOI: 10.1111/j.1365-2818.1894.tb00029.x.
  • Michalski J, Ratajczak E, Wiśniewski J. 1985. Roztocze (Acarina: Mesostigmata) towarzyszące kornikom (Coleptera: Scolytidae) Gór Świętokrzyskich. Prace Komisji Nauk Rolniczych i Komisji Nauk Leśnych 60:85–92.
  • Miko L, Stanko M. 1991. Small mammals as carriers of non-parasitic mites (Oribatida, Uropodina). In: Dusbábek F, Bukva V, editors. Modern acarology. Vol. 1. The Hague: Academia, Prague and SPB Academic Publishing bv. pp. 395–402.
  • Monzenga Lokela JC, Le Goff GJ, Kayisu K, Hance T. 2021. Phoretic mites associated with Rhynchophorus phoenicis Fabricius (1880) (Coleoptera: Curculionidae) in the Kisangani region, D. R Congo Acarologia 61(2):291–296. DOI: 10.24349/acarologia/20214431.
  • Moraza ML, Fernández M, Jurc M. 2013. Phoretic mites of the six-spined engraver beetle, Ips sexdentatus (Böerner, 1776) (Coleoptera, Scolytinae), on Pinus halepensis in Slovenia. International Journal of Acarology 39(8):597–599. DOI: 10.1080/01647954.2013.867900.
  • Morton A, García-Del-Pino F. 2011. Possible interaction of the phoretic mite Centrouropoda almerodai on the control of Rhynchophorus ferrugineus by entomopathogenic nematodes. IOBC-WPRS Bulletin. 66:363–366.
  • Moser JC. 1976a. Phoretic carrying capacity of flying southern pine beetles (Coleoptera: Scolytidae). The Canadian Entomologist 108(8):807–808. DOI: 10.4039/ent108807-8.
  • Moser JC. 1976b. Surveying mites (Acarina) phoretic on the southern pine beetle (Coleoptera: Scolytidae) with sticky traps. The Canadian Entomologist 108(8):809–813. DOI: 10.4039/Ent108809-8.
  • Moser JC, Blomquist SR. 2011. Phoretic arthropods of the red imported fire ant in Central Louisiana. Annals of the Entomological Society of America 104(5):886–894. DOI: 10.1603/AN11010.
  • Moser JC, Bogenschütz H. 1984. A key to the mites associated with flying Ips typographus in South Germany. Zeitschrift für Angewandte Entomologie 97(1–5):437–450. DOI: 10.1111/j.1439-0418.1984.tb03774.x.
  • Moser JC, Eidmann HH, Regnander JR. 1989. The mites associated with Ips typographus in Sweden. Annales Entomologici Fennici 55:23–27.
  • Moser JC, Konrad H, Blomquist SR, Kirisits T. 2010. Do mites phoretic on elm bark beetles contribute to the transmission of Dutch elm disease? Naturwissenschaften 97:219–227. DOI: 10.1007/s00114-009-0630-x.
  • Moser JC, Konrad H, Kirisits T, Carta LK. 2005. Phoretic mites and nematode associates of Scolytus multistriatus and Scolytus pygmaeus (Coleoptera: Scolytidae) in Austria. Agricultural and Forest Entomology 7(2):169–177. DOI: 10.1111/j.1461-9555.2005.00261.x.
  • Moser JC, Perry TJ, Furuta K. 1997. Phoretic mites and their hyperphoretic fungi associated with flying Ips typographus japonicus Niijima (Col., Scolytidae) in Japan. Journal of Applied Entomology 121(1–5):425–428. DOI: 10.1111/j.1439-0418.1997.tb01429.x.
  • Moser JC, Roton LM. 1971. Mites associated with southern pine bark beetles in Allen Parish, Louisiana. The Canadian Entomologist 103(12):1775–1798. DOI: 10.4039/Ent1031775-12.
  • Müller OF. 1776. Zoologiae Danicae Prodromus, seu Animalium Daniae et Norvegicae Indigenarum. Characteres, Nomina, et Synonyma Imprimis Popularum. Hafniae, Copenhagen. p. 282. DOI: 10.5962/bhl.title.13268.
  • Napierała A, Błoszyk J. 2013. Unstable microhabitats (merocenoses) as specific habitats of Uropodina mites (Acari: Mesostigmata). Experimental and Applied Acarology 60(2):163–180. DOI: 10.1007/s10493-013-9659-9.
  • Napierała A, Książkiewicz Z, Leśniewska M, Gwiazdowicz DJ, Mądra A, Błoszyk J. 2015. Phoretic relationships between uropodid mites (Acari: Mesostigmata) and centipedes (Chilopoda) in urban agglomeration areas. International Journal of Acarology 41(4):250–258. DOI: 10.1080/01647954.2015.1029968.
  • Noriega JA, Halliday B, Weston P, Thotagamuwa A, Gurr G. 2022. Beyond phoresy: Symbioses between dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) and mites (Acari). Entomologia Generalis 42(4):499–513. DOI: 10.1127/entomologia/2021/1406.
  • O’Donnell AE, Axtell RC. 1965. Predation by Fuscuropoda vegetans (Acarina: Uropodidae) on the house fly (Musca domestica). Annals of the Entomological Society of America 58(3):403–404. DOI: 10.1093/aesa/58.3.403.
  • O’Donnell AE, Nelson EL. 1967. Predation by Fuscuropoda vegetans (Acarina: Uropodidae) and Macrocheles muscaedomesticae (Acarina: Macrochelidae) on the eggs of the little house fly, Fannia canicularis. Journal of the Kansas Entomological Society 40(3):441–443.
  • Oudemans AC. 1902. New list of Dutch Acari. Second part. Tijdschrift voor Entomologie 45:1–52 + plates 1–6.
  • Oudemans AC. 1915. Acarologische Aanteekeningen. LVI. Entomologische berichten 4(83):180–188.
  • Oudemans AC. 1916. Acarologische aanteekeningen. LX. Entomologische berichten 4(91):307–316. DOI: 10.5962/bhl.part.1130.
  • Paraschiv M, Isaia G. 2020. Disparity of phoresy in mesostigmatid mites upon their specific carrier Ips typographus (Coleoptera: Scolytinae). Insects 11(11):771. DOI: 10.3390/insects11110771.
  • Paraschiv M, Martínez‑Ruiz C, Fernández MM. 2018. Dynamic associations between Ips sexdentatus (Coleoptera: Scolytinae) and its phoretic mites in a Pinus pinaster forest in northwest Spain. Experimental & Applied Acarology 75(3):369–381. DOI: 10.1007/s10493-018-0272-9.
  • Penttinen R, Viiri H, Moser JC. 2013. The mites (Acari) associated with bark beetles in the Koli National Park in Finland. Acarologia 53(1):3–15. DOI: 10.1051/acarologia/20132074.
  • Pereira MCSDA, Kontschán J, Bernardi LFDO, Hermes MG. 2016. Description of a new species of Oplitis Berlese, 1884, and the first record of a phoretic uropodid mite (Acari: Oplitidae) on a Neotropical solitary wasp (Hymenoptera: Vespidae). Zootaxa 4067(3):394–400. DOI: 10.11646/zootaxa.4067.3.8.
  • Pérez-Lachaud G, Klompen H, Poteaux C, Santamaría C, Armbrecht I, Beugnon G, Lachaud J-P. 2019. Context dependent life-history shift in Macrodinychus sellnicki mites attacking a native ant host in Colombia. Scientific Reports 9(1):1–10. DOI: 10.1038/s41598-019-44791-2.
  • Pernek M, Hrasovec B, Matosevic D, Pilas I, Kirisits T, Moser JC. 2008. Phoretic mites of three bark beetles (Pityokteines spp.) on silver fir. Journal of Pest Science 81(1):35–42. DOI: 10.1007/s10340-007-0182-9.
  • Pernek M, Wirth S, Blomquist SR, Avtzis DN, Moser JC. 2012. New associations of phoretic mites on Pityokteines curvidens (Coleoptera, Curculionidae, Scolytinae). Central European Journal Biology 7(1):63–68. DOI: 10.2478/s11535-011-0096-7.
  • Perotti MA, Braig HR. 2009. Phoretic mites associated with animal and human decomposition. Experimental and Applied Acarology 49(1–2):85–124. DOI: 10.1007/s10493-009-9280-0.
  • Pfammatter JA, Raffa KF. 2015. Do phoretic mites influence the reproductive success of Ips grandicollis (Coleoptera: Curculionidae)? Environmental Entomology 44(6):1498–1511. DOI: 10.1093/ee/nvv125.
  • Pfammatter JA, Moser JC, Raffa KF. 2013. Mites phoretic on Ips pini (Coleoptera: Curculionidae: Scolytinae) in Wisconsin red pine stands. Annals of the Entomological Society of America 106(2):204–213. DOI: 10.1603/AN12122.
  • Pfammatter JA, Coyle DR, Gandhi KJK, Hernandez N, Hofstetter RW, Moser JC, Raffa KF. 2016a. Structure of phoretic mites assemblages across subcortical beetle species at a regional scale. Environmental Entomology 45(1):53–65. DOI: 10.1093/ee/nvv150.
  • Pfammatter JA, Malas KM, Raffa KF. 2016b. Behaviours of phoretic mites (Acari) associated with Ips pini and Ips grandicollis (Coleoptera: Curculionidae) during host-tree colonization. Agricultural and Forest Entomology 18(2):108–118. DOI: 10.1111/afe.12143.
  • Philips JR. 2009. The mite (Acarina) fauna of trogid beetles (Coleoptera: Trogidae). International Journal of Acarology 35(1):1–17. DOI: 10.1080/01647950802709843.
  • Phillipsen WJ, Coppel HC. 1978. Uroobovella formosana sp. n. associated with the Formosan subterranean termite, Coptotermes formosanus Shiraki (Acarina: Uropodidae—Isoptera: Rhinotermitidae). Journal of the Kansas Entomological Society 51(1):22–27.
  • Poinar GO, Ćurčić BPM, Cokendolpher JC. 1998. Arthropod phoresy involving pseudoscorpion in the past and present. Acta Arachnologica 47(2):79–96. DOI: 10.2476/asjaa.47.79.
  • Porcelli F, Ragusa E, D’Onghia AM, Mizzi S, Mifsud D. 2009. Occurrence of Centrouropoda almerodai and Uroobovella marginata (Acari: Uropodina) phoretic on the Red Palm Weevil in Malta. Bulletin of the Entomological Society of Malta 2:61–66.
  • Pugh PJA, Llewellyn PJ, Robinson K, Shackley SE. 1997. The associations of phoretic mites (Acarina: Chelicerata) with sand-hoppers (Amphipoda: Crustacea) on the South Wales coast. Journal of Zoology 243(2):305–318. DOI: 10.1111/j.1469-7998.1997.tb02784.x.
  • Ragusa E, Porcelli F, Di Palma A, Ragusa Di Chiara S. 2008. Una specie di acaro associata al Punteruolo rosso delle palme: Centrouropoda almerodai (Uroactiininae, Uropodina). Regione Siciliana—Assessorato Agricoltura e Foreste Dipartimento Interventi Infrastrutturali—Servizi allo Sviluppo, editor La ricerca scientifica sul Punteruolo rosso e gli altri fitofagi delle palme in Sicili 1:79–82.
  • Ramsey GW. 1967. Phoretic mites associated with insects. New Zealand Entomologist 3(5):6–7. DOI: 10.1080/00779962.1966.9722853.
  • Raut SK, Panigrahi A. 1991. The mite Fuscuropoda marginata (C. L. Koch) for the control of pest slugs Laevicaulis alte (Ferussac). In: Dusbábek F, Bukva V, editors. Modern acarology. Vol. 2. The Hague: Academia, Prague and SPB Academic Publishing bv. pp. 683–687.
  • Rigby MC. 1996. Association of a juvenile phoretic uropodid mite with the beach hopper Traskorchestia traskiana (Stimpson, 1857) (Crustacea: Talitridae). Journal of Natural History 30(11):1617–1624. DOI: 10.1080/00222939600770941.
  • Rodrigueiro TSC, Do Prado AP. 2004. Macrocheles muscaedomesticae (Acari, Macrochelidae) and a species of Uroseius (Acari, Polyaspididae) phoretic on Musca domestica (Diptera, Muscidae): effects on dispersal and colonization of poultry manure. Iheringia Série Zoologia 94(2):181–185. DOI: 10.1590/S0073-47212004000200011.
  • Rubink WL, Delfinado-Baker M, Wilson WT, de Lourdes Gonzales-Gracia M, Gonzales-Cortés S. 1991. A phoretic uropodid mite associated with honey bee swarms of Northeastern Mexico. International Journal of Acarology 17(4):259–263. DOI: 10.1080/01647959108683916.
  • Schwan TG, Corwin D. 1987. Uropodid mites phoretic on fleas of ground squirrels in California. Proceedings of the Entomological Society of Washington. 89:790–793.
  • Schwarz HH, Starrach M, Koulianos S. 1998. Host specificity and permanence of associations between mesostigmatic mites (Acari: Anactinotrichida) and burying beetles (Coleoptera: Silphidae: Nicrophorus). Journal of Natural History 32(2):159–172. DOI: 10.1080/00222939800770101.
  • Seeman OD, Walter DE. 2023. Phoresy and mites: More than just a free ride. Annual Review of Entomology 68:69–88. DOI: 10.1146/annurev-ento-120220-013329.
  • Sellnick M. 1926. Alguns novos Acaros (Uropodidae) myrmecophilos e termitophilos. Archivos do Museu Nacional Rio de Janeiro 26:29–56.
  • Sellnick M. 1958. Eine neue Milbe aus Fliedermauskot (sic) in einer Höhle Südafrikas (Acarina-Uropodina). Journal of the Entomological Society of Southern Africa 21(2):274–285.
  • Slimane-Kharrat S, Ouali O. 2019. Mites associated with the red palm weevil (Rhynchophorus ferrugineus) in Tunisia. Tunisian Journal of Plant Protection 14(2):29–38.
  • Springett BP. 1968. Aspects of the relationship between burying beetles, Necrophorus spp. and the mite Poecilochirus necrophori Vitz. Journal of Animal Ecology 37(2):417–424. DOI: 10.2307/2957.
  • Stewart TB, Davis R. 1967. Notes on mites associated with coprophagous beetles. Journal of the Georgia Entomological Society 2:21–26.
  • Trouessart E, Berlese A. 1919. Generi nuovi di Acari. Redia 14:4.
  • Vissa S, Hofstetter RW, Bonifácio L, Khaustov A, Knee W, Uhey DA. 2019. Phoretic mite communities associated with bark beetles in the maritime and stone pine forests of Setúbal, Portugal. Experimental and Applied Acarology 77(2):117–131. DOI: 10.1007/s10493-019-00348-6.
  • Vissa S, Soderberg DN, Hofstetter RW. 2021. Field translocation of mountain pine beetles suggests phoretic mite communities are locally adapted, and mite populations respond variably to climate warming. Insects 12(2):131. DOI: 10.3390/insects12020131.
  • Vitzthum H. 1921. Acarologische Beobachtungen. 4. Reihe Archiv für Naturgeschichte 86(A, 10):1–69.
  • Vitzthum H. 1923. Acarologische Beobachtungen. 7 Reihe. Kommensalen der Ipiden. Archiv für Naturgeschichte 89(A2, 2):97–181.
  • Vitzthum H. 1926. Acari als Commensalen von Ipiden (Der Acarologischen Beobachtungen 11 Reihe). Zoologische Jahrbücher Abteilung für Systematik, Geographie und Biologie der Tiere 52:407–503.
  • Vrabec M, Kalúz S, Ferenčík J. 2012. Foretické roztoče na lykožrútovi smrekovom (Ips typographus) na vybraných lokalitách vo Vysokých Tatrách. Entomofauna carpathica 24(1):1–14.
  • Wiśniewski J. 1979a. Für die Fauna Polens neue Uropodina (Acari: Parasitiformes). Fragmenta Faunistica 25(5):73–76. DOI: 10.3161/00159301FF1979.25.5.073.
  • Wiśniewski J. 1979b. Gangsystematik der Parasitiformes. Teil 339. Zur Kenntnis der Uropodiden-Fauna Polens. Acarologie Schiftenreihe für vergleichende Milbenkunde 26:68–74.
  • Wiśniewski J. 1982. Für die Fauna Polens neue Uropodina (Acari: Parasitiformes). Teil. II. Fragmenta Faunistica 27(10):143–147. DOI: 10.3161/00159301FF1982.27.10.143.
  • Wiśniewski J, Hirschmann W. 1991. Ergänzungsangaben über Verbreitung und Lebensräume der Trichouropoda-Arten (Acarina, Uropodina). Fragmenta Faunistica 35(11):173–177. DOI: 10.3161/00159301FF1991.35.11.173.
  • Wiśniewski J, Hirschmann W. 1992. Neue Uropodina-Arten [Acarina] aus dem Staatlichen Museum fur Naturkunde in Lwow [Ukraine]. Bulletin of the Polish Academy of Sciences. Biological Sciences 41(1):43–65.
  • Wiśniewski J, Hirschmann W. 1993a. Gangsystematik der Parasitiformes. Teil 548. Katalog der Ganggattungen, Untergattungen, Gruppen und Arten der Uropodiden der Erde (Taxonomie, Literatur, Grösse, Verbreitung, Vorkommen). Acarologie Schriftenreihe für Vergleichende Milbenkunde 40:1–220.
  • Wiśniewski J, Hirschmann W. 1993b. Stadien von drei neuen Uroseius (Apionoseius)-Arten (Acarina, Uropodina) auf Trox-Arten (Coleoptera, Tenebrionidae) aus USA und Brasilien. Bulletin of the Polish Academy of Sciences Biological Sciences 41(1):85–97.
  • Wiśniewski J, Michalski J. 1984. Gangsystematik der Parasitiformes. Teil 456. Stadien von 3 neuen Trichouropoda-Arten aus Sibirien, nebst einigen zoogeographischen Angaben über die Uropodiden-Fauna der Sowjetunion (Trichouropodini, Uropodinae). Acarologie Schiftenreihe für vergleichende Milbenkunde 31:75–80.
  • Wiśniewski J, Hiramatsu N, Hirschmann W. 1992a. Gangsystematik der Parasitiformes. Teil 539. Gang und Stadien von 9 neuen Uroobovella-Arten der ipidis-Gruppe (Dinychini, Uropodinae). Acarologie Schiftenreihe für vergleichende Milbenkunde 39:109–128.
  • Wiśniewski J, Hirschmann W, Hiramatsu N. 1992b. Neue Centrouropoda-Arten (Uroactiniinae, Uropodina) aus den Philippinen, aus Brasilien und Mittelafrika. Acarologia 33(4):313–320.
  • Yasui Y. 2023. Mite dilemma: molting to acquire sexual maturity or not molting to ensure durability and dispersal ability in Phorytocarpais fimetorum (Parasitiformes; Gamasida; Parasitidae). Journal of Ethology 41(2):177–184. DOI: 10.1007/s10164-023-00783-4.
  • Zach P, Kršiak B, Kulfan J, Parák M, Kontschán J. 2016. Mites Trichouropoda and Uroobovella spp. (Uropodoidea) phoretic on bark beetles (Scolytinae): A comparison from a declining mountain spruce forest in Central Europe. International Journal of Acarology 42(4):212–217. DOI: 10.1080/01647954.2016.1154107.