Anatomy and relationships of the Middle Permian varanopid Heleosaurus scholtzi based on a social aggregation from the Karoo Basin of South Africa

LITERATURE CITED

• Anderson , J. S. and Reisz , R. R. 2004 . Pyozia mesenensis, a new, small varanopid (Synapsida, Eupelycosauria) from Russia: “pelycosaur” diversity in the Middle Permian. . Journal of Vertebrate Paleontology , 24 : 173 – 179 .
   Web of Science ®Google Scholar
• Berman , D. S. and Reisz , R. R. 1982 . Restudy of Mycterosaurus longiceps (Reptilia, Pelycosauria) from the Lower Permian of Texas. . Annals of Carnegie Museum , 51 : 423 – 453 .
   Google Scholar
• Botha-Brink , J. and Modesto , S. P. 2007 . A mixed-age classed ‘pelycosaur’ aggregation from South Africa: earliest evidence of parental care in amniotes? . Proceedings of the Royal Society B , 274 : 2829 – 2834 .
   PubMed Web of Science ®Google Scholar
• Broom , R. 1907 . On some new fossil reptiles from the Karroo beds of Victoria West, South Africa. . Transactions of the South African Philosophical Society , 18 : 31 – 42 .
   Google Scholar
• Broom , R. 1937 . A further contribution to our knowledge of the fossil reptiles of the Karroo. . Proceedings of the Zoological Society of London , 1937 : 299 – 318 .
   Google Scholar
• Carroll , R. L. 1976 . “ Eosuchians and the origin of archosaurs; ” . In Essays on Palaeontology in Honour of Loris Shano Russell Edited by: Churcher , C. S. 58 – 79 . Toronto : Royal Ontario Museum .
   Google Scholar
• Dias-da-Silva , S. , Modesto , S. P. and Schultz , C. L. 2007 . New material of Procolophon (Parareptilia: Procolophonoidea) from the Lower Triassic of Brazil, with remarks on the ages of the Sanga do Cabral and Buena Vista formations of South America. . Canadian Journal of Earth Sciences , 43 : 1685 – 1693 .
   Web of Science ®Google Scholar
• Dilkes , D. W. and Reisz , R. R. 1996 . First record of a basal synapsid (‘mammal-like reptile’) in Gondwana. Proceedings of the Royal Society of London . Series B , 263 : 1165 – 1170 .
   Google Scholar
• Gauthier , J. A. , Kluge , A. G. and Rowe , T. 1988 . Amniote phylogeny and the importance of fossils. . Cladistics , 4 : 105 – 209 .
   PubMed Web of Science ®Google Scholar
• Kemp , T. S. 1982 . Mammal-like Reptiles and the Origin of Mammals 363 London : Academic Press .
   Google Scholar
• Kitching , J. W. 1977 . The distribution of the Karroo vertebrate fauna. Bernard Price Institute for Palaeontological Research, University of the Witwatersrand . Memoir , 1 : 1 – 131 .
   Google Scholar
• Langston , J. W. and Reisz , R. R. 1981 . Aerosaurus wellesi, new species, a varanopseid mammal-like reptile (Synapsida: Pelycosauria) from the Lower Permian of New Mexico. . Journal of Vertebrate Paleontology , 1 : 73 – 96 .
   Google Scholar
• Lucas , S. G. 2006 . “ Global Permian tetrapod biostratigraphy and biochronology; ” . In Non-Marine Permian Biostratigraphy and Biochronology Edited by: Lucas , S. G. , Cassinis , G. and Schneider , J. W. 65 – 93 . Geological Society of London, Special Publication . 265
   Google Scholar
• Maddin , H. C. , Evans , D. C. and Reisz , R. R. 2006 . An Early Permian varanodontine varanopid (Synapsida: Eupelycosauria) from the Richards Spur Locality, Oklahoma. . Journal of Vertebrate Paleontology , 26 : 957 – 966 .
   Web of Science ®Google Scholar
• Modesto , S. P. , Sidor , C. A. , Rubidge , B. S. and Welman , J. 2001 . A second varanopseid skull from the Upper Permian of South Africa: implications for Late Permian ‘pelycosaur’ evolution. . Lethaia , 34 : 249 – 259 .
   Web of Science ®Google Scholar
• Osborn , H. F. 1903 . On the primary division of the Reptilia into two subclasses, Synapsida and Diapsida. . Science , 17 : 275 – 276 .
   PubMedGoogle Scholar
• Piñeiro , G. , Verde , M. , Ubilla , M. and Ferigolo , J. 2003 . First basal synapsids (“pelycosaurs”) from the Upper Permian-?Lower Triassic of Uruguay, South America. . Journal of Paleontology , 77 : 389 – 392 .
   Web of Science ®Google Scholar
• Reisz , R. R. and Berman , D. S. 2001 . The skull of Mesenosaurus romeri, a small varanopseid (Synapsida: Eupelycosauria) from the Upper Permian of the Mezen River Basin, northern Russia. . Annals of Carnegie Museum , 70 : 113 – 132 .
   Web of Science ®Google Scholar
• Reisz , R. R. and Dilkes , D. W. 2003 . Archaeovenator hamiltonensis, a new varanopid (Synapsida: Eupelycosauria) from the Upper Carboniferous of Kansas. . Canadian Journal of Earth Sciences , 40 : 667 – 678 .
   Web of Science ®Google Scholar
• Reisz , R. R. and Laurin , M. 2004 . A reevaluation of the enigmatic Permian synapsid Watongia and of its stratigraphic significance. . Canadian Journal of Earth Sciences , 41 : 377 – 386 .
   Web of Science ®Google Scholar
• Reisz , R. R. and Modesto , S. P. 2007 . Heleosaurus scholtzi from the Permian of South Africa: a varanopid synapsid, not a diapsid reptile. . Journal of Vertebrate Paleontology , 27 : 734 – 739 .
   Web of Science ®Google Scholar
• Reisz , R. R. , Dilkes , D. W. and Berman , D. S. 1998 . Anatomy and relationships of Elliotsmithia longiceps Broom, a small synapsid (Eupelycosauria: Varanopseidae) from the Late Permian of South Africa. . Journal of Vertebrate Paleontology , 18 : 602 – 611 .
   Web of Science ®Google Scholar
• Romer , A. S. and Price , L. I. 1940 . Review of the Pelycosauria. Geological Society of America . Special Papers , 40 : 1 – 538 .
   Google Scholar
• Smith , R. M. H. and Keyser , A. W. 1995 . “ Biostratigraphy of the Tapinocephalus Assemblage Zone; ” . In Biostratigraphy of the Beaufort Group (Karoo Supergroup) Edited by: Rubidge , B. S. 8 – 12 . South African Committee for Stratigraphy Biostratigraphic Series . 1
   Google Scholar
• Swofford , D. L. 2002 . PAUP: Phylogenetic analysis Using Parsimony (and Other Methods). Version 4.0b8 Massachusetts : Sinauer Associates: Sunderland .
   Google Scholar